Lista de Anfíbios

Brazilian Amphibians: List of Species

Magno V. Segalla1,*, Ulisses Caramaschi2, Carlos Alberto Gonalves Cruz2, Taran Grant3, Clio F. B. Haddad4, Jos A. Langone5, Paulo Christiano de Anchietta Garcia6

1 Laboratrio de Herpetologia. Museu de Histria Natural Capo da Imbuia, CEP82810-080, Curitiba, PR, Brazil.2 Universidade Federal do Rio de Janeiro, Museu Nacional, Departamento de Vertebrados, Quinta da Boa Vista, So Cristvo, CEP20940-040, Rio de Janeiro, RJ, Brazil.3 Departamento de Zoologia, Instituto de Biocincias, Universidade de So Paulo, CEP05508-090, So Paulo, SP, Brazil.4 Departamento de Zoologia, Instituto de Biocincias, Universidade Estadual Paulista, Caixa Postal 199, CEP13506-906, Rio Claro, SP, Brazil.5 Departamento de Herpetologa, Museo Nacional de Historia Natural, Casilla de Correo 399, 11.000, Montevideo, Uruguay.6 Departamento de Zoologia, Instituto de Cincias Biolgicas; Universidade Federal de Minas Gerais, Avenida Antnio Carlos, 6627, Pampulha, CEP31270-901,

Belo Horizonte, MG, Brazil.* Corresponding author. Email: [email protected]

16. Allobates masniger (Morales, 2002 2000)17. Allobates nidicola (Caldwell & Lima, 2003)18. Allobates olfersioides (A. Lutz, 1925)19. Allobates paleovarzensis Lima, Caldwell, Biavati & Mon-

tanarin, 201020. Allobates subfolionidificans (Lima, Sanchez & Souza, 2007)21. Allobates sumtuosus (Morales, 2002 2000)22. Allobates vanzolinius (Morales, 2002 2000)

Family Aromobatidae (Allobatinae)

23. Anomaloglossus baeobatrachus (Boistel & de Massari, 1999)

24. Anomaloglossus roraima (La Marca, 1998)25. Anomaloglossus stepheni (Martins, 1989)26. Anomaloglossus tamacuarensis (Myers & Donelly, 1997)

Family Brachycephalidae

27. Brachycephalus alipioi Pombal & Gasparini, 200628. Brachycephalus atelopoide Miranda-Ribeiro, 192029. Brachycephalus brunneus Ribeiro, Alves, Haddad & dos

Reis, 200530. Brachycephalus bufonoides Miranda-Ribeiro, 192031. Brachycephalus crispus Condez, Clemente-Carvalho &

Haddad, 201432. Brachycephalus didactylus (Izecksohn, 1971)33. Brachycephalus ephippium (Spix, 1824)34. Brachycephalus ferruginus Alves, Ribeiro, Haddad & dos

Reis, 200635. Brachycephalus garbeanus Miranda-Ribeiro, 192036. Brachycephalus guarani Clemente-Carvalho, Giaretta,

Condez, Haddad & dos Reis, 201237. Brachycephalus hermogenesi (Giaretta & Sawaya, 1998)38. Brachycephalus izecksohni Ribeiro, Alves, Haddad & dos

Reis, 200539. Brachycephalus margaritatus Pombal & Izecksohn, 201140. Brachycephalus nodoterga Miranda-Ribeiro, 192041. Brachycephalus pernix Pombal, Wistuba & Bornschein,

1998

The following list includes all recognized species of amphi-bians known to occur within the political borders of Brazil. The taxonomy follows Frost (2014). The present list differs from previous ones by incuding subfamilies (in parentheses follo-wing familes).

The known amphibian fauna of Brazil comprises 1026 spe-cies. The vast majority of species are anurans, including 988 species representing 19 families and 87 genera, followed by caecilians, with 33 species in four families and 12 genera, and salamanders, with five species in a single family and genus.

ORDER ANURA

Family Allophrynidae

1. Allophryne ruthveni Gaige, 19262. Allophryne relicta Caramaschi, Orrico, Faivovich, Dias &

Sol, 2013

Family Alsodidae

3. Limnomedusa macroglossa (Dumril & Bibron, 1841)

Family Aromobatidae (Allobatinae)

4. Allobates brunneus (Cope, 1887)5. Allobates caeruleodactylus (Lima & Caldwell, 2001)6. Allobates conspicuus (Morales, 2002 2000)7. Allobates crombiei (Morales, 2002 2000)8. Allobates femoralis (Boulenger, 1884 1883)9. Allobates flaviventris Melo-Sampaio, Souza & Peloso,

201310. Allobates fuscellus (Morales, 2002 2000)11. Allobates gasconi (Morales, 2002 2000)12. Allobates grillisimilis Simes, Sturaro, Peloso & Lima,

201313. Allobates goianus (Bokermann, 1975)14. Allobates hodli Simes, Lima & Farias, 201015. Allobates marchesianus (Melin, 1941)

Herpetologia Brasileira - Volume 3 - Nmero 2 - Julho de 2014

Mudanas Taxonmicas 37

42. Brachycephalus pitanga Alves, Sawaya, dos Reis & Ha-ddad, 2009

43. Brachycephalus pombali Alves, Ribeiro, Haddad & dos Reis, 2006

44. Brachycephalus pulex Napoli, Caramaschi, Cruz & Dias, 2011

45. Brachycephalus toby Haddad, Alves, Clemente-Carvalho & Reis, 2010

46. Brachycephalus tridactylus Garey, Lima, Hartmann & Haddad, 2012

47. Brachycephalus vertebralis Pombal, 200148. Ischnocnema bdita Canedo & Pimenta, 201049. Ischnocnema bolbodactyla (A. Lutz, 1925)50. Ischnocnema concolor Targino, Costa & Carvalho e Silva,

200951. Ischnocnema epipeda (Heyer, 1984)52. Ischnocnema erythromera (Heyer, 1984)53. Ischnocnema gehrti (Miranda-Ribeiro, 1926)54. Ischnocnema gualteri (B. Lutz, 1974)55. Ischnocnema guentheri (Steindachner, 1864)56. Ischnocnema henselii (Peters, 1872)57. Ischnocnema hoehnei (B. Lutz, 1959 1958)58. Ischnocnema holti (Cochran, 1948)59. Ischnocnema izecksohni (Caramaschi & Kisteumacher,

1989 1988)60. Ischnocnema juipoca (Sazima & Cardoso, 1978)61. Ischnocnema karst Canedo, Targino, Leite & Haddad, 201262. Ischnocnema lactea (Miranda-Ribeiro, 1923)63. Ischnocnema manezinho (Garcia, 1996)64. Ischnocnema melanopygia Targino, Costa & Carvalho e

Silva, 200965. Ischnocnema nanahallux Brusquetti, Thom, Canedo,

Condez, Haddad, 201366. Ischnocnema nasuta (A. Lutz, 1925)67. Ischnocnema nigriventris (A. Lutz, 1925)68. Ischnocnema octavioi (Bokermann, 1965)69. Ischnocnema oea (Heyer, 1984)70. Ischnocnema paranaensis (Langone & Segalla, 1996)71. Ischnocnema parva (Girard, 1853)72. Ischnocnema penaxavantinho Giaretta, Toffoli & Oliveira,

200773. Ischnocnema pusilla (Bokermann, 1967)74. Ischnocnema randorum (Heyer, 1985)75. Ischnocnema sambaqui (Castanho & Haddad, 2000)76. Ischnocnema spanios (Heyer, 1985)77. Ischnocnema surda Canedo, Pimenta, Leite & Caramas-

chi, 201078. Ischnocnema venancioi (B. Lutz, 1959 1958)79. Ischnocnema verrucosa (Reinhardt & Ltken, 1862)80. Ischnocnema vizottoi Martins & Haddad, 2010

Family Bufonidae

81. Amazophrynella bokermanni (Izecksohn, 1994 1993)82. Amazophrynella manaos Rojas, Carvalho, Gordo, vila,

Farias & Hrbek, 2014

83. Amazophrynella minuta (Melin, 1941)84. Amazophrynella vote Avila, Carvalho, Gordo, Kawashita-

-Ribeiro & Morais, 201285. Atelopus flavescens Dumril & Bibron, 184186. Atelopus hoogmoedi Lescure, 197487. Atelopus spumarius Cope, 187188. Dendrophryniscus berthalutzae Izecksohn, 1994 199389. Dendrophryniscus brevipollicatus Jimnez de la Espada,

1871 187090. Dendrophryniscus carvalhoi Izecksohn, 1994 199391. Dendrophryniscus krausae Cruz & Fusinatto, 200892. Dendrophryniscus leucomystax Izecksohn, 196893. Dendrophryniscus oreites Recoder, Teixeira, Cassimiro,

Camacho & Rodrigues, 201094. Dendrophryniscus organensis Carvalho-e-Silva, Mongin,

Izecksohn & Carvalho-e-Silva, 201095. Dendrophryniscus proboscideus (Boulenger, 1882)96. Dendrophryniscus skuki (Caramaschi, 2012)97. Dendrophryniscus stawiarskyi Izecksohn, 1994 199398. Frostius erythrophthalmus Pimenta & Caramaschi, 200799. Frostius pernambucensis (Bokermann, 1962)100. Melanophryniscus admirabilis Di Bernardo, Maneyro &

Grillo, 2006101. Melanophryniscus alipioi Langone, Segalla, Bornschein &

de S, 2008102. Melanophryniscus atroluteus (Miranda-Ribeiro, 1920)103. Melanophryniscus cambaraensis Braun & Braun, 1979104. Melanophryniscus devincenzii Klappenbach, 1968105. Melanophryniscus dorsalis (Mertens, 1933)106. Melanophryniscus fulvoguttatus (Mertens, 1937)107. Melanophryniscus klappenbachi Prigioni & Langone,

2000108. Melanophryniscus macrogranulosus Braun, 1973109. Melanophryniscus montevidensis (Philippi, 1902)110. Melanophryniscus moreirae (Miranda-Ribeiro, 1920)111. Melanophryniscus pachyrhynus (Miranda-Ribeiro, 1920)112. Melanophryniscus peritus Carmaschi & Cruz, 2011113. Melanophryniscus sanmartini Klappenbach, 1968114. Melanophryniscus setiba Peloso, Faivovich, Grant, Gaspa-

rini & Haddad, 2012115. Melanophryniscus simplex Caramaschi & Cruz, 2002116. Melanophryniscus spectabilis Caramaschi & Cruz, 2002117. Melanophryniscus tumifrons (Boulenger, 1905)118. Melanophryniscus vilavelhensis Steinback-Padilha, 2009119. Oreophrynella quelchii Boulenger, 1895120. Oreophrynella weiassipuensis Searis, Nascimento &

Villarreal, 2005121. Rhaebo ecuadorensis Mueses-Cisneros, Cisneros-Heredia

& Mcdiarmid, 2012122. Rhaebo guttatus (Schneider, 1799)123. Rhinella abei (Baldissera-Jr,Caramaschi & Haddad,

2004)124. Rhinella achavali (Maneyro, Arrieta & de S, 2004)125. Rhinella acutirostris (Spix, 1824)126. Rhinella arenarum (Hensel, 1867)127. Rhinella azarai (Gallardo, 1965)128. Rhinella bergi (Cspedez, 2000 1999)


Mudanas Taxonmicas38

129. Rhinella castaneotica (Caldwell, 1991)130. Rhinella ceratophrys (Boulenger, 1882)131. Rhinella cerradensis Maciel, Brando, Campos & Sebben,

2007132. Rhinella crucifer (Wied-Neuwied, 1821)133. Rhinella dapsilis (Myers & Carvalho, 1945)134. Rhinella dorbignyi (Dumril & Bibron, 1841)135. Rhinella fernandezae (Gallardo, 1957)136. Rhinella granulosa (Spix, 1824)137. Rhinella henseli (A. Lutz, 1934)138. Rhinella hoogmoedi Caramaschi & Pombal, 2006139. Rhinella icterica (Spix, 1824)140. Rhinella inopina Vaz-Silva, Valdujo & Pombal, 2012141. Rhinella jimi (Stevaux, 2002)142. Rhinella magnussoni Lima, Menin & Arajo, 2007143. Rhinella major (Muller & Helmich, 1936)144. Rhinella margaritifera (Laurenti, 1768 )145. Rhinella marina (Linnaeus, 1758)146. Rhinella martyi Fouquet, Gaucher, Blanc & Vlez-Rodri-

guez, 2007147. Rhinella merianae Gallardo, 1965148. Rhinella mirandaribeiroi (Gallardo, 1965)149. Rhinella nattereri Bokermann, 1967150. Rhinella ocellata (Gnther, 1859 1858)151. Rhinella ornata (Spix, 1824)152. Rhinella paraguayensis vila, Pansonato & Strssmann,

2010153. Rhinella proboscidea (Spix, 1824)154. Rhinella pygmaea (Myers & Carvalho, 1952)155. Rhinella roqueana (Melin, 1941)156. Rhinella rubescens (A. Lutz, 1925)157. Rhinella schneideri (Werner, 1894)158. Rhinella scitula (Caramaschi & Niemeyer, 2003)159. Rhinella veredas (Brando,Maciel & Sebben, 2007)

Family Centrolenidae (Centroleninae)

160. Cochranella adenocheira Harvey & Noonan, 2005 Incer-tae sedis

161. Teratohyla midas (Lynch & Duellman, 1973)162. Vitreorana eurygnatha (A. Lutz, 1925)163. Vitreorana parvula (Boulenger, 1895 1894)164. Vitreorana ritae (B. Lutz in B. Lutz & Kloss, 1952)165. Vitreorana uranoscopa (Mller, 1924)

Family Centrolenidae (Hyalinobatrachinae)

166. Hyalinobatrachium cappellei (van Lidth de Jeude, 1904)167. Hyalinobatrachium carlesvilai Castroviejo-Fisher, Padial,

Chaparro, Aguayo & De la Riva, 2009168. Hyalinobatrachium iaspidiense (Ayarzaqena, 1992)169. Hyalinobatrachium mondolfii Searis & Ayarzaguena,

2001170. Hyalinonatrachium munozorum (Lynch & Duellman,

1973)

Family Ceratophryidae

171. Ceratophrys aurita (Raddi, 1823)172. Ceratophrys cornuta (Linnaeus, 1758)173. Ceratophrys cranwelli Barrio, 1980174. Ceratophrys joazeirensis Mercadal de Barrio, 1986175. Ceratophrys ornata (Bell, 1843)176. Lepidobatrachus asper (Budgett, 1899)

Family Craugastoridae (Craugastorinae)

177. Haddadus aramunha (Cassimiro, Verdade & Rodrigues, 2008)

178. Haddadus binotatus (Spix, 1824)179. Haddadus plicifer (Boulenger, 1888)180. Strabomantis sulcatus (Cope, 1874)

Family Craugastoridae (Holoadeninae)

181. Eleutherodactylus bilineatus (Bokermann, 1975) Incer-tae sedis

182. Barycholos ternetzi (Miranda Ribeiro, 1937)183. Euparkerella brasiliensis (Parker, 1926)184. Euparkerella cochranae Izecksohn, 1988185. Euparkerella robusta Izecksohn, 1988186. Euparkerella tridactyla Izecksohn, 1988187. Holoaden bradei B. Lutz, 1959 1958188. Holoaden luederwaldti Miranda-Ribeiro, 1920189. Holoaden pholeter Pombal, Siqueira, Dorigo, Vrcibradic &

Rocha, 2008190. Holoaden suarezi Martins & Zaher, 2013191. Noblella myrmecoides (Lynch, 1976)

Family Craugastoridae (Ceuthomantinae)

192. Ceuthomantis cavernibardus (Myers & Donnelly, 1997)193. Oreobates crepitans (Bokermann, 1965)194. Oreobates heterodactylus (Miranda-Ribeiro, 1937)195. Oreobates quixensis Jimnez de la Espada, 1872196. Oreobates remotus Teixeira, Amaro, Recoder, Sena &

Rodrigues, 2012197. Pristimantis academicus Lehr, Moravec & Urrutia, 2010198. Pristimantis achuar Elmer & Cannatella, 2008199. Pristimantis acuminatus (Schreve, 1935)200. Pristimantis altamazonicus (Barbour & Dunn, 1921)201. Pristimantis aureolineatus Guayasamin, Ron, Cisneros-

-Heredia, Lamar & McCracken, 2006202. Pristimantis aureoventris Kok, Means & Bossuyt, 2011203. Pristimantis buccinator (Rodriguez, 1994)204. Pristimantis carvalhoi (B. Lutz in B. Lutz & Kloss, 1952)205. Pristimantis chiastonotus (Lynch & Hoogmoed, 1977)206. Pristimantis conspicillatus (Gnther, 1858)207. Pristimantis delius (Duellman & Mendelson, 1995)208. Pristimantis diadematus (Jimnez de la Espada, 1875)



209. Pristimantis dundeei (Heyer & Muoz, 1999)210. Pristimantis eurydactylus (Hedges & Schlter, 1992)211. Pristimantis fenestratus (Steindachner, 1864)212. Pristimantis gutturalis (Hoogmoed, Lynch & Lescure,

1977)213. Pristimantis inguinalis (Parker, 1940)214. Pristimantis lacrimosus (Jimnez de la Espada, 1875)215. Pristimantis lanthanites (Lynch, 1975)216. Pristimantis malkini (Lynch, 1980)217. Pristimantis marmoratus (Boulenger, 1900)218. Pristimantis martiae (Lynch, 1974)219. Pristimantis memorans (Myers & Donelly, 1997)220. Pristimantis ockendeni (Boulenger, 1912)221. Pristimantis orcus Lehr, Catenazzi & Rodriguez, 2009222. Pristimantis paulodutrai (Bokermann, 1975 1974)223. Pristimantis peruvianus (Melin, 1941)224. Pristimantis ramagii (Boulenger, 1888)225. Pristimantis reichlei Padial & de La Riva, 2009226. Pristimantis skydmainos (Flores & Rodriguez, 1997)227. Pristimantis toftae (Duellman, 1978)228. Pristimantis variabilis (Lynch, 1968)229. Pristimantis ventrigranulosus Maciel, Vaz-Silva, de Oli-

veira & Padial, 2012230. Pristimantis ventrimarmoratus (Boulenger, 1912)231. Pristimantis vilarsi (Melin, 1941)232. Pristimantis vinhai (Bokermann, 1975)233. Pristimantis zeuctotylus (Lynch & Hoogmoed, 1977)234. Pristimantis zimmermanae (Heyer & Hardy, 1991)

Family Cycloramphidae

235. Cycloramphus acangatan Verdade & Rodrigues, 2003236. Cycloramphus asper Werner, 1899237. Cycloramphus bandeirensis Heyer, 1983238. Cycloramphus bolitoglossus (Werner, 1897)239. Cycloramphus boraceiensis Heyer, 1983240. Cycloramphus brasiliensis (Steindachner, 1864)241. Cycloramphus carvalhoi Heyer, 1983242. Cycloramphus catarinensis Heyer, 1983243. Cycloramphus cedrensis Heyer, 1983244. Cycloramphus diringshofeni Bokermann, 1957245. Cycloramphus dubius (Miranda-Ribeiro, 1920)246. Cycloramphus duseni (Andersson, 1914)247. Cycloramphus eleutherodactylus (Miranda-Ribeiro,

1920)248. Cycloramphus faustoi Brasileiro, Haddad, Sawaya & Sazi-

ma, 2007249. Cycloramphus fuliginosus Tschudi, 1838250. Cycloramphus granulosus A. Lutz, 1929251. Cycloramphus izecksohni Heyer, 1983252. Cycloramphus juimirim Haddad & Sazima, 1989253. Cycloramphus lithomimeticus DaSilva & Ouvernay 2012254. Cycloramphus lutzorum Heyer, 1983255. Cycloramphus migueli Heyer, 1988256. Cycloramphus mirandaribeiroi Heyer, 1983257. Cycloramphus ohausi (Wandolleck, 1907)



258. Cycloramphus organensis Weber, Verdade, Salles, Fouquet & Carvalho-e-Silva, 2011

259. Cycloramphus rhyakonastes Heyer, 1983260. Cycloramphus semipalmatus (Miranda-Ribeiro, 1920)261. Cycloramphus stejnegeri (Noble, 1924)262. Cycloramphus valae Heyer, 1983263. Thoropa lutzi Cochran, 1938264. Thoropa megatympanum Caramaschi & Sazima, 1984265. Thoropa miliaris (Spix, 1824)266. Thoropa petropolitana (Wandolleck, 1907)267. Thoropa saxatilis Crocoft & Heyer, 1988268. Thoropa taophora (Miranda-Ribeiro, 1923)269. Zachaenus carvalhoi Izecksohn, 19831982270. Zachaenus parvulus (Girard, 1853)

Family Dendrobatidae (Colostethinae)

271. Ameerega berohoka Vaz-Silva & Maciel, 2011272. Ameerega braccata (Steindachner, 1864)273. Ameerega flavopicta (A. Lutz, 1925)274. Ameerega hahneli (Boulenger, 1884 1883)275. Ameerega macero (Rodriguez & Myers, 1993)276. Ameerega petersi (Silverstone, 1976)277. Ameerega picta (Bibron in Tschudi, 1838)278. Ameerega pulchripecta (Silverstone, 1976)279. Ameerega trivittata (Spix, 1824)

Family Dendrobatidae (Dendrobatinae)

280. Adelphobates castaneoticus (Caldwell & Myers, 1990)281. Adelphobates galactonotus (Steindachner, 1864)282. Adelphobates quinquevittatus (Steindachner, 1864)283. Dendrobates leucomelas Steindachner, 1864284. Dendrobates tinctorius (Cuvier, 1797)285. Ranitomeya amazonica (Schulte, 1999)286. Ranitomeya cyanovittata Perez-Pea, Chavez, Twomey &

Brown, 2010287. Ranitomeya defleri Twomey & Brown, 2009288. Ranitomeya flavovittata (Schulte, 1999)289. Ranitomeya sirensis (Aichinger, 1991)290. Ranitomeya toraro Brown, Caldwell, Twomey, Melo-Sam-

paio & Souza, 2011291. Ranitomeya uakarii Brown, Schulte & Summers, 2006292. Ranitomeya vanzolinii (Myers, 1982)293. Ranitomeya variabilis (Zimmermann & Zimmermann,

1988)294. Ranitomeya yavaricola Perez-Pea, Chavez, Twomey &

Brown, 2010

Family Dendrobatidae (Hyloxalinae)

295. Hyloxalus chlorocraspedus (Caldwell, 2005)

Family Eleutherodactylidae (Phyzelaphryninae)

296. Adelophryne adiastola Hoogmoed & Lescure, 1984297. Adelophryne baturitensis Hoogmoed, Borges & Cascon,

1994298. Adelophryne gutturosa Hoogmoed & Lescure, 1984299. Adelophryne maranguapensis Hoogmoed, Borges & Cas-

con, 1994300. Adelophryne meridionalis Santana, Fonseca, Neves &

Carvalho 2013 2012301. Adelophryne mucronatus Loureno-de-Moraes, Sol &

Toledo, 2012302. Adelophryne pachydactyla Hoogmoed, Borges & Cascon,

1994303. Phyzelaphryne miriamae Heyer, 1977

Family Hemiphractidae

304. Fritziana fissilis (Miranda Ribeiro, 1920)305. Fritiziana goeldii (Boulenger, 1895 1894)306. Fritziana ohausi (Wandolleck, 1907)307. Gastrotheca albolineata (Lutz & Lutz, 1939)308. Gastrotheca ernestoi Miranda Ribeiro, 1920309. Gastrotheca fissipes (Boulenger, 1888)310. Gastrotheca flamma Junc & Nunes, 2008311. Gastrotheca fulvorufa (Andersson, 1911)312. Gastrotheca megacephala Izecksohn, Carvalho-e-Silva &

Peixoto, 2009313. Gastrotheca microdiscus (Andersson in Lnnberg & An-

dersson, 1910)314. Gastrotheca prasina Teixeira, Dal Vechio, Recoder, Carna-

val, Strangas, Damasceno, de Sena & Rodrigues, 2012315. Gastrotheca pulchra Caramaschi & Rodrigues, 2007316. Gastrotheca recava Teixeira, Dal Vechio, Recoder, Carna-

val, Strangas, Damasceno, de Sena & Rodrigues, 2012317. Hemiphractus helioi Sheil & Mendelson, 2001318. Hemiphractus scutatus (Spix, 1824)319. Stefania neblinae Carvalho, MacCulloch, Bonora & Vogt,

2010320. Stefania tamacuarina Myers & Donnelly, 1997

Family Hylidae (Hylinae)

321. Hyla imitator (Barbour & Dunn, 1921) Incertae sedis322. Calamita melanorabdotus sensu Frost, 2006 Hyla mela-

norabdota (Schneider, 1799) Incertae sedis323. Aparasphenodon arapap Pimenta, Napoli & Haddad,

2009324. Aparasphenodon bokermanni Pombal, 1993325. Aparasphenodon brunoi Miranda-Ribeiro, 1920326. Aparasphenodon venezolanus (Mertens, 1950)327. Aparasphenodon pomba Assis, Santana, Da Silva, Quinte-

la & Feio, 2013328. Aplastodiscus albofrenatus (A. Lutz, 1924)329. Aplastodiscus albosignatus (A. Lutz & B. Lutz, 1938)

330. Aplastodiscus arildae (Cruz & Peixoto, 1987 1985)331. Aplastodiscus callipygius (Cruz & Peixoto, 1985 1984)332. Aplastodiscus cavicola (Cruz & Peixoto, 1985 1984)333. Aplastodiscus cochranae (Mertens, 1952)334. Aplastodiscus ehrhardti (Mller, 1924)335. Aplastodiscus eugenioi (Carvalho e Silva & Carvalho e

Silva, 2005)336. Aplastodiscus flumineus (Cruz & Peixoto, 1985 1984)337. Aplastodiscus ibirapitanga (Cruz, Pimenta & Silvano,

2003)338. Aplastodiscus leucopygius (Cruz & Peixoto, 1985 1984)339. Aplastodiscus musicus (B. Lutz, 1948)340. Aplastodiscus perviridis A. Lutz in B. Lutz, 1950341. Aplastodiscus sibilatus (Cruz, Pimenta & Silvano, 2003)342. Aplastodiscus weygoldti (Cruz & Peixoto, 1987 1985)343. Bokermannohyla ahenea (Napoli & Caramaschi, 2004)344. Bokermannohyla alvarengai (Bokermann, 1956)345. Bokermannohyla astartea (Bokermann, 1977)346. Bokermannohyla caramaschii (Napoli, 2005)347. Bokermannohyla carvalhoi (Peixoto, 1981)348. Bokermannohyla capra Napoli & Pimenta, 2009349. Bokermannohyla caramaschii (Napoli, 2005)350. Bokermannohyla circumdata (Cope, 1871)351. Bokermannohyla claresignata (A. Lutz & B. Lutz, 1939)352. Bokermannohyla clepsydra (A. Lutz, 1925)353. Bokermannohyla diamantina Napoli & Junc, 2006354. Bokermannohyla feioi (Napoli & Caramaschi, 2004)355. Bokermannohyla flavopicta Leite, Pezzuti & Garcia,

2012356. Bokermannohyla gouveai (Peixoto & Cruz, 1992)357. Bokermannohyla hylax (Heyer, 1985)358. Bokermannohyla ibitiguara (Cardoso, 1983)359. Bokermannohyla ibitipoca (Caramaschi & Feio, 1990)360. Bokermannohyla itapoty Lugli & Haddad, 2006361. Bokermannohyla izecksohni (Jim & Caramaschi, 1979)362. Bokermannohyla juiju Faivovich, Luigli, Loureno &

Haddad, 2009363. Bokermannohyla langei (Bokermann, 1965)364. Bokermannohyla lucianae (Napoli & Pimenta, 2003)365. Bokermannohyla luctuosa (Pombal & Haddad, 1993)366. Bokermannohyla martinsi (Bokermann, 1964)367. Bokermannohyla nanuzae (Bokermann & Sazima, 1973)368. Bokermannohyla napolii Carvalho, Giaretta & Magrini,

2012369. Bokermannohyla oxente Lugli & Haddad, 2006370. Bokermannohyla pseudopseudis (Miranda-Ribeiro, 1937)371. Bokermannohyla ravida (Caramaschi, Napoli & Bernar-

des, 2001)372. Bokermannohyla sagarana Leite, Pezzuti & Drummond,

2011373. Bokermannohyla Sapiranga Brando, Magalhes, Garda,

Campos, Sebben & Maciel, 2012374. Bokermannohyla saxicola (Bokermann, 1964)375. Bokermannohyla sazimai (Cardoso & Andrade,

19831982)376. Bokermannohyla vulcaniae (Vasconcelos & Giaretta, 2004

2003)



377. Corythomantis galeata Pombal, Menezes, Fontes, Nunes, Rocha & Van Sluys, 2012

378. Corythomantis greeningi Boulenger, 1896379. Dendropsophus acreanus (Bokermann, 1964)380. Dendropsophus anataliasiasi (Bokermann, 1972)381. Dendropsophus anceps (A. Lutz, 1929)382. Dendropsophus araguaya (Napoli & Caramaschi, 1998)383. Dendropsophus berthalutzae (Bokermann, 1962)384. Dendropsophus bifurcus (Andersson, 1945)385. Dendropsophus bipunctatus (Spix, 1824)386. Dendropsophus bokermanni (Goin, 1960)387. Dendropsophus branneri (Cochran, 1948)388. Dendropsophus brevifrons (Duellman & Crump, 1974)389. Dendropsophus cachimbo (Napoli & Caramaschi, 1999)390. Dendropsophus cerradensis (Napoli & Caramaschi, 1998)391. Dendropsophus cruzi (Pombal & Bastos, 1998)392. Dendropsophus decipiens (A. Lutz, 1925)393. Dendropsophus dutrai (Gomes & Peixoto, 1996)394. Dendropsophus elegans (Wied-Neuwied, 1824)395. Dendropsophus elianeae (Napoli & Caramaschi, 2000)396. Dendropsophus gaucheri (Lescure & Marty, 2000)397. Dendropsophus giesleri (Mertens, 1950)398. Dendropsophus haddadi (Bastos & Pombal, 1996)399. Dendropsophus haraldschultzi (Bokermann, 1962)400. Dendropsophus jimi (Napoli & Caramaschi, 1999)401. Dendropsophus koechlini (Duellman & Trueb, 1989)402. Dendropsophus leali (Bokermann, 1964) 403. Dendropsophus leucophyllatus (Beireis, 1783)404. Dendropsophus limai (Bokermann, 1962)405. Dendropsophus marmoratus (Laurenti, 1768406. Dendropsophus melanargyreus (Cope, 1887)407. Dendropsophus meridianus (B. Lutz, 1954)408. Dendropsophus microcephalus (Cope, 1886)409. Dendropsophus microps (Peter, 1872)410. Dendropsophus minimus (Ahl, 1933)411. Dendropsophus minusculus (Rivero, 1971)412. Dendropsophus minutus (Peters, 1872)413. Dendropsophus miyatai (Vigle & Goberdhan-Vigle, 1990)414. Dendropsophus nahdereri (B. Lutz & Bokermann, 1963)415. Dendropsophus nanus (Boulenger, 1889)416. Dendropsophus novaisi (Bokermann, 1968)417. Dendropsophus oliveirai (Bokermann, 1963)418. Dendropsophus parviceps (Boulenger, 1882)419. Dendropsophus pauiniensis (Heyer, 1977)420. Dendropsophus pseudomeridianus (Cruz, Caramaschi &

Dias, 2000)421. Dendropsophus rhea (Napoli & Caramaschi, 1999)422. Dendropsophus rhodopeplus (Gnther, 1859 1858)423. Dendropsophus riveroi (Cochran & Goin, 1970)424. Dendropsophus rossalleni (Goin, 1959)425. Dendropsophus rubicundulus (Reinhardt & Ltken,

18621861)426. Dendropsophus ruschii (Weygoldt & Peixoto, 1987)427. Dendropsophus sanborni (Schmidt, 1944)428. Dendropsophus sarayacuensis (Shreve, 1935)429. Dendropsophus schubarti (Bokermann, 1963)430. Dendropsophus seniculus (Cope, 1868)

431. Dendropsophus soaresi (Caramaschi & Jim, 1983)432. Dendropsophus studerae (Carvalho e Silva, Carvalho e

Silva & Izecksohn, 2003)433. Dendropsophus timbeba (Martins & Cardoso, 1987)434. Dendropsophus tintinnabulum (Melin, 1941)435. Dendropsophus triangulum (Gnther, 1869 1868)436. Dendropsophus tritaeniatus (Bokermann, 1965)437. Dendropsophus walfordi (Bokermann, 1962)438. Dendropsophus werneri (Cochran, 1952)439. Dendropsophus xapuriensis (Martins & Cardoso, 1987)440. Dryaderces inframaculatus (Boulenger, 1882)441. Dryaderces pearsoni (Gaige, 1929)442. Ecnomiohyla tuberculosa (Boulenger, 1882)443. Hypsiboas albomarginatus (Spix, 1824)444. Hypsiboas albopunctatus (Spix, 1824)445. Hypsiboas atlanticus (Caramaschi & Velosa, 1996)446. Hypsiboas bandeirantes Caramaschi & Cruz, 2013447. Hypsiboas beckeri (Caramaschi & Cruz, 2004)448. Hypsiboas benitezi (Rivero, 1961)449. Hypsiboas bischoffi (Boulenger, 1887)450. Hypsiboas boans (Linnaeus, 1758)451. Hypsiboas botumirim Caramaschi, Cruz & Nascimento,

2009452. Hypsiboas buriti (Caramaschi & Cruz, 1999)453. Hypsiboas caingua (Carrizo, 1991 1990)454. Hypsiboas caipora Antunes, Faivovich & Haddad, 2008455. Hypsiboas calcaratus (Troschel in Schomburgk, 1848)456. Hypsiboas cinerascens (Spix, 1824)457. Hypsiboas cipoensis (B. Lutz, 1968)458. Hypsiboas crepitans (Wied-Neuwied, 1824)459. Hypsiboas curupi Garcia, Faivovichi & Haddad, 2007460. Hypsiboas cymbalum (Bokerman, 1963)461. Hypsiboas dentei (Bokermann, 1967)462. Hypsiboas ericae (Caramaschi & Cruz, 2000)463. Hypsiboas exastis (Caramaschi & Rodriguez, 2003)464. Hypsiboas faber (Wied-Neuwied, 1821)465. Hypsiboas fasciatus (Gnther, 1859 1858)466. Hypsiboas freicanecae (Carnaval & Peixoto, 2004)467. Hypsiboas geographicus (Spix, 1824)468. Hypsiboas goianus (B. Lutz, 1968)469. Hypsiboas guentheri (Boulenger, 1886)470. Hypsiboas jaguariaivensis Caramaschi, Cruz & Segalla,

2010471. Hypsiboas joaquini (Lutz, 1968)472. Hypsiboas lanciformis (Cope, 1871)473. Hypsiboas latistriatus (Caramaschi & Cruz, 2004)474. Hypsiboas leptolineatus (P. Braun & C. Braun, 1977)475. Hypsiboas leucocheilus (Carmaschi & Niemeyer, 2003)476. Hypsiboas lundii (Burmeister, 1856)477. Hypsiboas marginatus (Boulenger, 1887)478. Hypsiboas microderma (Pyburn, 1977)479. Hypsiboas multifasciatus (Gnther, 18591858)480. Hypsiboas nympha Faivovich, Moravec, Cisneros-heredia

& Khler, 2006481. Hypsiboas ornatissimus (Noble, 1923)482. Hypsiboas pardalis (Spix, 1824)483. Hypsiboas paranaiba Carvalho, Giaretta & Facure, 2010



484. Hypsiboas phaeopleura (Caramaschi & Cruz, 2000)485. Hypsiboas poaju Garcia, Peixoto & Haddad, 2008486. Hypsiboas polytaenius (Cope, 1870 1869)487. Hypsiboas pombali (Caramaschi, Pimenta & Feio, 2004)488. Hypsiboas prasinus (Burmeister, 1856)489. Hypsiboas pulchellus (Dumril & Bibron, 1841)490. Hypsiboas punctatus (Schneider, 1799)491. Hypsiboas raniceps Cope, 1862492. Hypsiboas secedens (B. Lutz, 1963)493. Hypsiboas semiguttatus (A. Lutz, 1925)494. Hypsiboas semilineatus (Spix, 1824)495. Hypsiboas stellae Kwet, 2008496. Hypsiboas stenocephalus (Caramaschi & Cruz, 1999)497. Hypsiboas tepunianus Barrio-Amoros & Brewer-Carias,

2008498. Hypsiboas wavrini (Parker, 1936)499. Itapotihyla langsdorffii (Dumril & Bibron, 1841)500. Lysapsus boliviana Gallardo, 1961501. Lysapsus caraya Gallardo, 1964502. Lysapsus laevis (Parker, 1935)503. Lysapsus limellum Cope, 1862504. Osteocephalus buckleyi (Boulenger, 1882)505. Osteocephalus cabrerai (Cochran & Goin, 1970)506. Osteocephalus helenae (Ruthven, 1919)507. Osteocephalus leprieurii (Dumril & Bibron, 1841)508. Osteocephalus oophagus Jungfer & Schiesari, 1995509. Osteocephalus planiceps Cope, 1874510. Osteocephalus subtilis Martins & Cardoso, 1987511. Osteocephalus taurinus Steindachner, 1862512. Osteocephalus vilarsi (Melin, 1941)513. Phyllodytes acuminatus Bokermann, 1966514. Phyllodytes brevirostris Peixoto & Cruz, 1988515. Phyllodytes edelmoi Peixoto, Caramaschi & Freire, 2003516. Phyllodytes gyrinaethes Peixoto, Caramaschi & Freire,

2003517. Phyllodytes kautskyi Peixoto & Cruz, 1988518. Phyllodytes luteolus Wied-Neuwied, 1824519. Phyllodytes maculosus Cruz, Feio & Cardoso, 2006

2007520. Phyllodytes melanomystax Caramaschi, Da Silva & Britto-

-Pereira, 1992521. Phyllodytes punctatus Caramaschi & Peixoto, 2004522. Phyllodytes tuberculosus Bokermann, 1966523. Phyllodytes wuchereri (Peters, 1873 1872)524. Pseudis bolbodactyla A. Lutz, 1925525. Pseudis cardosoi Kwet, 2000526. Pseudis fusca Garman, 1883527. Pseudis minuta Gnther, 1858528. Pseudis paradoxa (Linnaeus, 1758)529. Pseudis platensis Gallardo, 1961530. Pseudis tocantins Caramaschi & Cruz, 1998531. Scarthyla goinorum (Bokermann, 1962)532. Scinax acuminatus (Cope, 1862)533. Scinax agilis (Cruz & Peixoto, 1983)534. Scinax albicans (Bokermann, 1967)535. Scinax alcatraz (B. Lutz, 1973)536. Scinax alter (B. Lutz, 1973)

537. Scinax angrensis (B. Lutz, 1973)538. Scinax arduous Peixoto, 2002539. Scinax argyreornatus (Miranda-Ribeiro, 1926)540. Scinax ariadne (B. Lutz, 1973)541. Scinax aromothyella Faivovich, 2005542. Scinax atratus (Peixoto, 1989)543. Scinax auratus (Wied-Neuwied, 1821)544. Scinax baumgardneri (Rivero, 1961)545. Scinax belloni Faivoivch, Gasparini & Haddad, 2010546. Scinax berthae (Barrio, 1962)547. Scinax blairi (Fouquette & Pyburn, 1972)548. Scinax boesemani (Goin, 1966)549. Scinax brieni (Witte, 1930)550. Scinax cabralensis Drummond, Bata & Pires, 2007551. Scinax caldarum (B. Lutz, 1968)552. Scinax camposseabrai (Bokermann, 1968)553. Scinax canastrensis (Cardoso & Haddad, 1982)554. Scinax cardosoi (Carvalho e Silva & Peixoto, 1991)555. Scinax carnevallii (Caramaschi & Kisteumacher, 1989)556. Scinax catharinae (Boulenger, 1888)557. Scinax centralis Pombal & Bastos, 1996558. Scinax constrictus Lima, Bastos & Giaretta, 2004559. Scinax cosenzai Lacerda, Peixoto & Feio, 2012560. Scinax cretatus Nunes & Pombal, 2011561. Scinax crospedospilus (A. Lutz, 1925)562. Scinax cruentommus (Duellman, 1972)563. Scinax curicica Pugliesse, Pombal & Sazima, 2004564. Scinax cuspidatus (A. Lutz, 1925)565. Scinax dolloi (Werner, 1903)566. Scinax duartei (B. Lutz, 1951)567. Scinax eurydice (Bokermann, 1968)568. Scinax exiguus (Duellman, 1986)569. Scinax faivovichi Brasileiro, Oyamaguchi & Haddad,

2007570. Scinax flavoguttatus (Lutz & Lutz, 1939) Lutz, 1939)571. Scinax funereus (Cope, 1874)572. Scinax fuscomarginatus (A. Lutz, 1925)573. Scinax fuscovarius (A. Lutz, 1925)574. Scinax garbei (Miranda-Ribeiro, 1926)575. Scinax granulatus (Peters, 1871)576. Scinax hayii (Barbour, 1909)577. Scinax heyeri (Peixoto & Weygoldt, 1986)578. Scinax hiemalis (Haddad & Pombal, 1987)579. Scinax humilis (B. Lutz, 1954)580. Scinax imbegue Nunes, Kwet & Pombal, 2012581. Scinax inesperatus Da Silva & Alves-Silva, 2011582. Scinax juncae Nunes & Pombal, 2010583. Scinax jureia (Pombal & Gordo, 1991)584. Scinax kautskyi (Carvalho e Silva & Peixoto, 1991)585. Scinax lindsayi Pyburn, 1992586. Scinax littoralis (Pombal & Gordo, 1991)587. Scinax littoreus (Peixoto, 1988)588. Scinax longilineus (B. Lutz, 1968)589. Scinax luizotavioi (Caramaschi & Kisteumacher, 1989)590. Scinax machadoi (Bokermann & Sazima, 1973)591. Scinax madeirae Brusquetti, Jansen, Barrio-Amors,

Segalla & Haddad, 2014



592. Scinax maracaya (Cardoso & Sazima, 1980)593. Scinax melloi (Peixoto, 1989)594. Scinax muriciensis Cruz, Nunes & Lima, 2011595. Scinax nasicus (Cope, 1862)596. Scinax nebulosus (Spix, 1824)597. Scinax obtriangulatus (B. Lutz, 1973)598. Scinax pachycrus (Miranda-Ribeiro, 1937)599. Scinax pedromedinae (Henle, 1991)600. Scinax peixotoi Brasileiro, Haddad, Sawaya & Martins,

2007601. Scinax perereca Pombal, Haddad & Kasahara, 1995602. Scinax perpusillus (A. Lutz & B. Lutz, 1939)603. Scinax pinima (Bokermann & Sazima, 1973)604. Scinax pombali Loureno, Carvalho, Baeta, Pezzuti & Lei-

te, 2013605. Scinax proboscideus (Brongersma, 1933)606. Scinax ranki (Andrade & Cardoso, 1987)607. Scinax rizibilis (Bokermann, 1964)608. Scinax rogerioi Pugliesi, Bata & Pombal, 2009609. Scinax rostratus (Peter, 1863)610. Scinax ruber (Laurenti, 1768)611. Scinax satermawe Sturaro & Peloso, 2014612. Scinax skaios Pombal, Carvalho, Canelas & Bastos, 2010613. Scinax similis (Cochran, 1952)614. Scinax skuki Lima, Cruz & Azevedo, 2011615. Scinax squalirostris (A. Lutz, 1925)616. Scinax strigilatus (Spix, 1824)617. Scinax tigrinus Nunes, Carvalho & Pereira, 2010618. Scinax tymbamirim Nunes, Kwet & Pombal, 2012619. Scinax trapicheiroi (B. Lutz, 1954)620. Scinax tupinamb Silva & Alves-Silva, 2008621. Scinax uruguayus (Schmidt, 1944)622. Scinax v-signatus (B. Lutz, 1968)623. Scinax villasboasi Brusquetti, Jansen, Barrio-Amors,

Segalla & Haddad, 2014624. Scinax x-signatus (Spix, 1824)625. Sphaenorhynchus botocudo Caramaschi, Almeida & Gas-

parini, 2009626. Sphaenorhynchus mirim Caramaschi, Almeida & Gaspari-

ni, 2009627. Sphaenorhynchus bromelicola Bokermann, 1966628. Sphaenorhynchus caramaschii Toledo, Garcia, Lingnau &

Haddad, 2007629. Sphaenorhynchus carneus (Cope, 1868)630. Sphaenorhynchus dorisae (Goin, 1957)631. Sphaenorhynchus lacteus (Daudin, 1800)632. Sphaenorhynchus orophilus (A. Lutz & B. Lutz, 1938)633. Sphaenorhynchus palustris Bokermann, 1966634. Sphaenorhynchus pauloalvini Bokermann, 1973635. Sphaenorhynchus planicola (A. Lutz & B. Lutz, 1938)636. Sphaenorhynchus prasinus Bokermann, 1973637. Sphaenorhynchus surdus (Cochran, 1953)638. Tepuihyla exophthalma (Smith & Noonan, 2001)639. Trachycephalus atlas Bokermann, 1966640. Trachycephalus cunauaru Gordo, Toledo, Surez, Kawa-

shita-Ribeiro, vila, Morais & Nunes, 2013641. Trachycephalus coriaceus (Peters, 1867)

642. Trachycephalus dibernardoi Kwet & Sol, 2008643. Trachycephalus hadroceps (Duellman & Hoogmoed, 1992)644. Trachycephalus helioi Nunes, Surez, Gordo & Pombal,

2013645. Trachycephalus imitatrix (Miranda-Ribeiro, 1926)646. Trachycephalus lepidus (Pombal, Haddad & Cruz, 2003)647. Trachycephalus mambaiensis Cintra, Silva, Silva_Jr, Gar-

cia & Zaher, 2009648. Trachycephalus mesophaeus (Hensel, 1867)649. Trachycephalus nigromaculatus Tschudi, 1838650. Trachycephalus resinifictrix (Goeldi, 1907)651. Trachycephalus typhonius (Linnaeus, 1758)652. Xenohyla eugenioi Caramaschi, 1998653. Xenohyla truncata (Izecksohn, 1959)

Family Hylidae (Phyllomedusinae)

654. Agalychnis spera Peters, 1873 1872655. Agalychnis granulosa Cruz, 1989 1988656. Cruziohyla craspedopus (Funkhouser, 1957)657. Phasmahyla cochranae (Bokermann, 1966)658. Phasmahyla cruzi Carvalho e Silva, Silva & Carvalho e

Silva, 2009659. Phasmahyla exilis (Cruz, 1980)660. Phasmahyla guttata (A. Lutz, 1924)661. Phasmahyla jandaia (Bokermann & Sazima, 1978)662. Phasmahyla spectabilis Cruz, Feio & Nascimento, 2008663. Phasmahyla timb Cruz, Napoli & Fonseca, 2008664. Phrynomedusa appendiculata (Lutz, 1925)665. Phrynomedusa bokermanni Cruz, 1991666. Phrynomedusa fimbriata Miranda-Ribeiro, 1923667. Phrynomedusa marginata (Izecksohn & Cruz, 1976)668. Phrynomedusa vanzolinii Cruz, 1991669. Phyllomedusa atelopoides Duellman, Cadle & Cannatella,

1988670. Phyllomedusa ayeaye (B. Lutz, 1966)671. Phyllomedusa azurea Cope, 1862672. Phyllomedusa bahiana A. Lutz, 1925673. Phyllomedusa bicolor (Boddaert, 1772)674. Phyllomedusa boliviana Boulenger, 1902675. Phyllomedusa burmeisteri Boulenger, 1882676. Phyllomedusa camba De la Riva, 2000 1999677. Phyllomedusa centralis Bokermann, 1965678. Phyllomedusa distincta A. Lutz in B. Lutz, 1950679. Phyllomedusa hypochondrialis (Daudin, 1800)680. Phyllomedusa iheringii Boulenger, 1885681. Phyllomedusa megacephala (Miranda-Ribeiro, 1926)682. Phyllomedusa nordestina Caramaschi, 2006683. Phyllomedusa oreades Brando, 2002684. Phyllomedusa palliata Peters, 1873 1872685. Phyllomedusa rohdei Mertens, 1926686. Phyllomedusa sauvagii Boulenger, 1882687. Phyllomedusa tarsius (Cope, 1868)688. Phyllomedusa tetraploidea Pombal & Haddad, 1992689. Phyllomedusa tomopterna (Cope, 1868)690. Phyllomedusa vaillantii Boulenger, 1882



Family Hylodidae

691. Crossodactylus aeneus Mller, 1924692. Crossodactylus bokermanni Caramaschi & Sazima, 1985693. Crossodactylus boulengeri (De Witte, 1930)694. Crossodactylus caramaschii Bastos & Pombal, 1995695. Crossodactylus cyclospinus Nascimento, Cruz & Feio, 2005696. Crossodactylus dantei Carcerelli & Caramaschi, 1993

1992697. Crossodactylus dspar A. Lutz, 1925698. Crossodactylus gaudichaudii Dumril & Bibron, 1841699. Crossodactylus grandis B. Lutz, 1951700. Crossodactylus lutzorum Carcerelli & Caramaschi, 1993

1992701. Crossodactylus schmidti Gallardo, 1961702. Crossodactylus timbuhy Pimenta, Cruz & Caramaschi,

2014703. Crossodactylus trachystomus (Reinhardt & Ltken,

18621861)704. Crossodactylus werneri Pimenta, Cruz & Caramaschi, 2014705. Hylodes amnicola Pombal, Feio & Haddad, 2002706. Hylodes asper (Mller, 1924)707. Hylodes babax Heyer, 1982708. Hylodes cardosoi Lingnau, Canedo & Pombal, 2008709. Hylodes charadranaetes Heyer & Cocroft, 1986710. Hylodes dactylocinus Pavan, Narvaes & Rodrigues, 2001711. Hylodes fredi Canedo & Pombal, 2007712. Hylodes glaber (Miranda-Ribeiro, 1926)713. Hylodes heyeri Haddad, Pombal & Bastos, 1996714. Hylodes lateristrigatus (Baumann, 1912)715. Hylodes magalhaesi (Bokermann, 1964)716. Hylodes meridionalis (Mertens, 1927)717. Hylodes mertensi (Bokermann, 1956)718. Hylodes nasus (Lichtenstein, 1823)719. Hylodes ornatus (Bokermann, 1967)720. Hylodes otavioi Sazima & Bokermann, 1983 1982721. Hylodes perere Silva & Benmaman, 2008722. Hylodes perplicatus (Miranda-Ribeiro, 1926)723. Hylodes phyllodes Heyer & Cocroft, 1986724. Hylodes pipilans Canedo & Pombal, 2007725. Hylodes regius Gouva, 1979726. Hylodes sazimai Haddad & Pombal, 1995727. Hylodes uai Nascimento, Pombal & Haddad, 2001728. Hylodes vanzolinii Heyer, 1982729. Megaelosia apuana Pombal, Prado & Canedo, 2003730. Megaelosia bocainensis Giaretta, Bokermann & Haddad,

1993731. Megaelosia boticariana Giaretta & Aguiar, 1998732. Megaelosia goeldii (Baumann, 1912)733. Megaelosia jordanensis (Heyer, 1983)734. Megaelosia lutzae Izecksohn & Gouva, 1985735. Megaelosia massarti (Witte, 1930)

Family Leptodactylidae (Leiuperinae)

736. Edalorhina perezi Jimnez de la Espada, 1871 1870

737. Engystomops petersi Jimnez de la Espada, 1872738. Engystomops freibergi (Donoso-Barros, 1969)739. Physalaemus aguirrei Bokermann, 1966740. Physalaemus albifrons (Spix, 1824)741. Physalaemus albonotatus (Steindachner, 1864)742. Physalaemus angrensis Weber, Gonzaga & Carvalho e

Silva, 2005743. Physalaemus atlanticus Haddad & Sazima, 2004744. Physalaemus barrioi Bokermann, 1967745. Physalaemus biligonigerus (Cope, 1861 1860)746. Physalaemus bokermanni Cardoso & Haddad, 1985747. Physalaemus caete Pombal & Madureira, 1997748. Physalaemus Camacan Pimenta, Cruz & Silvano, 2005749. Physalaemus centralis Bokermann, 1962750. Physalaemus cicada Bokermann, 1966751. Physalaemus crombiei Heyer & Wolf, 1989752. Physalaemus cuvieri Fitzinger, 1826753. Physalaemus deimaticus Sazima & Caramaschi, 1988

1986754. Physalaemus ephippifer (Steindachner, 1864)755. Physalaemus erikae Cruz & Pimenta, 2004756. Physalaemus erythros Caramaschi, Feio & Guimares-

-Neto, 2003757. Physalaemus evangelistai Bokermann, 1967758. Physalaemus feioi Cassini, Cruz & Caramaschi, 2010759. Physalaemus gracilis (Boulenger, 1883)760. Physalaemus henselii (Peters, 1872)761. Physalaemus insperatus Cruz, Cassini & Caramaschi,

2008762. Physalaemus irroratus Cruz, Nascimento & Feio, 2007763. Physalaemus jordanensis Bokermann, 1967764. Physalaemus kroyeri (Reinhardt & Ltken, 1862 1861)765. Physalaemus lateristriga (Steindachner, 1864)766. Physalaemus lisei Braun & Braun, 1977767. Physalaemus maculiventris (Lutz, 1925)768. Physalaemus marmoratus (Reinhardt & Ltken, 1862

1861)769. Physalaemus maximus Feio, Pombal & Caramaschi, 1999770. Physalaemus moreirae (Miranda-Ribeiro, 1937)771. Physalaemus nanus (Boulenger, 1888)772. Physalaemus nattereri (Steindachner, 1863)773. Physalaemus obtectus Bokermann, 1966774. Physalaemus olfersii (Lichtenstein & Martens, 1856)775. Physalaemus orophilus Cassini, Cruz & Caramaschi, 2010776. Physalaemus riograndensis Milstead, 1960777. Physalaemus rupestris Caramaschi, Carcerelli & Feio,

1991778. Physalaemus signifer (Girard, 1853)779. Physalaemus soaresi Izecksohn, 1965780. Physalaemus spiniger (Miranda-Ribeiro, 1926)781. Pleurodema alium Maciel & Nunes, 2010782. Pleurodema bibroni Tschudi, 1838783. Pleurodema brachyops (Cope, 1869 1868)784. Pleurodema diplolister (Peters, 1870)785. Pseudopaludicola ameghini (Cope, 1887)786. Pseudopaludicola boliviana Parker, 1927787. Pseudopaludicola canga Giaretta & Kokubum, 2003



788. Pseudopaludicola ceratophryes Rivero & Serna, 1984789. Pseudopaludicola facureae Andrade & Carvalho, 2013790. Pseudopaludicola falcipes (Hensel, 1867)791. Pseudopaludicola giarettai Carvalho, 2012792. Pseudopaludicola hyleaustralis Pansonato, Morais, vila,

kawashita-Ribeiro, Strssmann & Martin, 2013793. Pseudopaludicola mineira Lobo, 1994794. Pseudopaludicola murundu Toledo, Siqueira, Duarte,

Veiga-Menoncello, Recco-Pimentel & Haddad, 2010795. Pseudopaludicola mystacalis (Cope, 1887)796. Pseudopaludicola parnaiba Roberto, Cardozo & Avila,

2013797. Pseudopaludicola pocoto Magalhes, Loebmann, Koku-

bum, Haddad & Garda, 2014798. Pseudopaludicola saltica (Cope, 1887)799. Pseudopaludicola serrana Toledo, 2010800. Pseudopaludicola ternetzi Miranda-Ribeiro, 1937

Family Leptodactylidae (Leptodactylinae)

801. Adenomera ajurauna (Berneck, Costa & Garcia 2008)802. Adenomera andreae (Mller, 1923)803. Adenomera araucria Kwet & Angulo, 2003804. Adenomera bokermanni (Heyer, 1973)805. Adenomera cotuba Carvalho & Giaretta, 2013806. Adenomera diptyx (Boettger, 1885)807. Adenomera engelsi Kwet, Steiner & Zillikens, 2009)808. Adenomera heyeri Boistel, Massary & Angulo, 2006809. Adenomera hylaedactyla (Cope, 1868)810. Adenomera juikitam Carvalho & Giaretta, 2013811. Adenomera marmorata (Steindachner, 1867)812. Adenomera martinezi (Bokermann, 1956)813. Adenomera nana Mller, 1922814. Adenomera saci Carvalho & Giaretta, 2013815. Adenomera thomei (Almeida & ngulo) 2006816. Hydrolaetare dantasi (Bokermann, 1959)817. Hydrolaetare schmidti (Cochran & Goin, 1959)818. Leptodactylus bolivianus Boulenger, 1898819. Leptodactylus bufonius Boulenger, 1894820. Leptodactylus caatingae Heyer & Junc, 2003821. Leptodactylus camaquara Sazima & Bokermann, 1978822. Leptodactylus chaquensis Cei, 1950823. Leptodactylus cunicularius Sazima & Bokermann,

1978824. Leptodactylus cupreus Caramaschi, Feio & So-Pedro,

2008825. Leptodactylus didymus Heyer, Garca-Lopez & Cardoso,

1996826. Leptodactylus diedrus Heyer, 1994827. Leptodactylus discodactylus Boulenger, 1884 1883828. Leptodactylus elenae Heyer, 1978829. Leptodactylus flavopictus Lutz, 1926830. Leptodactylus furnarius Sazima & Bokermann, 1978831. Leptodactylus fuscus (Schneider, 1799)832. Leptodactylus gracilis (Dumril & Bibron, 1841)833. Leptodactylus guianensis Heyer & De S, 2011

834. Leptodactylus hylodes (Reinhardt & Ltken, 1862 1861)

835. Leptodactylus jolyi Sazima & Bokermann, 1978836. Leptodactylus knudseni Heyer, 1972837. Leptodactylus labyrinthicus (Spix, 1824)838. Leptodactylus laticeps Boulenger, 1918839. Leptodactylus latinasus Jimnez de la Espada, 1875840. Leptodactylus latrans (Steffen, 1815)841. Leptodactylus lauramiriamae Heyer & Crombie, 2005842. Leptodactylus leptodactyloides (Andersson, 1945)843. Leptodactylus lineatus (Schneider, 1799)844. Leptodactylus longirostris Boulenger, 1882845. Leptodactylus macrosternum Miranda-Ribeiro, 1926846. Leptodactylus marambaiae Izecksohn, 1976847. Leptodactylus myersi Heyer, 1995848. Leptodactylus mystaceus (Spix, 1824)849. Leptodactylus mystacinus (Burmeister, 1861)850. Leptodactylus natalensis A. Lutz, 1930851. Leptodactylus notoaktites Heyer, 1978852. Leptodactylus ochraceus Lutz, 1930853. Leptodactylus oreomantis Carvalho, Leite & Pezzuti,

2013854. Leptodactylus paraensis Heyer, 2005855. Leptodactylus pentadactylus (Laurenti, 1768)856. Leptodactylus petersii (Steindachner, 1864)857. Leptodactylus plaumanni Ahl, 1936858. Leptodactylus podicipinus (Cope, 1862)859. Leptodactylus pustulatus (Peters, 1870)860. Leptodactylus rhodomystax Boulenger, 1884 1883861. Leptodactylus rhodonotus (Gnther, 1869 1868)862. Leptodactylus riveroi Heyer & Pyburn, 1983863. Leptodactylus rugosus Noble, 1923864. Leptodactylus sabanensis Heyer, 1994865. Leptodactylus sertanejo Giaretta & Costa, 2007866. Leptodactylus spixi Heyer, 1983867. Leptodactylus stenodema Jimnez de la Espada, 1875868. Leptodactylus syphax Bokermann, 1969869. Leptodactylus tapiti Sazima & Bokermann, 1978870. Leptodactylus troglodytes A. Lutz, 1926871. Leptodactylus validus Garman, 1888872. Leptodactylus vastus A. Lutz, 1930873. Leptodactylus viridis Jim & Spirandeli-Cruz, 1973874. Leptodactylus wagneri (Peters, 1862)

Family Leptodactylidae (Paratelmatobiinae)

875. Crossodactylodes bokermanni Peixoto, 1983 1982876. Crossodactylodes itambe Barata, Santos, Leite & Garcia,

2013877. Crossodactylodes izecksohni Peixoto, 1983 1982878. Crossodactylodes pintoi Cochran, 1938879. Crossodactylodes septentrionalis Teixeira, Recoder, Ama-

ro, Damasceno, Cassimiro & Rodrigues, 2013880. Paratelmatobius cardosoi Pombal & Haddad, 1999881. Paratelmatobius gaigeae (Cochran, 1938)882. Paratelmatobius lutzii Lutz & Carvalho, 1958



883. Paratelmatobius Mantiqueira Pombal & Haddad, 1999884. Paratelmatobius poecilogaster Giaretta & Castanho, 1990885. Paratelmatobius yepiranga Garcia, Berneck & Costa, 2009886. Rupirana cardosoi Heyer, 1999887. Scythrophrys sawayae (Cochran, 1953)

Family Microhylidae (Gastrophryninae)

888. Arcovomer passarellii Carvalho, 1954889. Chiasmocleis alagoanus Cruz, Caramaschi & Freire, 1999890. Chiasmocleis albopunctata (Boettger, 1885)891. Chiasmocleis antenori (Walker, 1973)892. Chiasmocleis atlantica Cruz, Caramaschi & Izecksohn,

1997893. Chiasmocleis avilapiresae Peloso & Sturaro 2008894. Chiasmocleis bassleri Dunn, 1949895. Chiasmocleis capixaba Cruz, Caramaschi & Izecksohn,

1997896. Chiasmocleis carvalhoi Cruz, Caramaschi & Izecksohn,

1997897. Chiasmocleis centralis Bokermann, 1952898. Chiasmocleis cordeiroi Caramaschi & Pimenta, 2003899. Chiasmocleis crucis Caramaschi & Pimenta, 2003900. Chiasmocleis gnoma Canedo, Dixo & Pombal, 2004901. Chiasmocleis haddadi Peloso, Sturaro, Forlani, Gaucher,

Motta & Wheeler, 2014902. Chiasmocleis hudsoni Parker, 1940903. Chiasmocleis leucosticta (Boulenger, 1888)904. Chiasmocleis Mantiqueira Cruz, Feio & Cassini, 2007905. Chiasmocleis mehelyi Caramaschi & Cruz, 1997906. Chiasmocleis papachibe Peloso, Sturaro, Forlani, Gaucher,

Motta & Wheeler, 2014907. Chiasmocleis quilombola Tonini, Forlani & de S, 2014908. Chiasmocleis royi Peloso, Sturaro, Forlani, Gaucher, Mot-

ta & Wheeler, 2014909. Chiasmocleis Sapiranga Cruz, Caramaschi & Napoli, 2007910. Chiasmocleis schubarti Bokermann, 1952911. Chiasmocleis shudikarensis Dunn, 1949912. Chiasmocleis tridactyla (Duellman & Medelson, 1995)913. Chiasmocleis ventrimaculata (Andersson, 1945)914. Ctenophryne geayi Mocquard, 1904915. Dasypops schirchi Miranda-Ribeiro, 1924916. Dermatonotus muelleri (Boettger, 1885)917. Elachistocleis bicolor (Valenciennes in Gurin-Menville,

1838)918. Elachistocleis bumbameuboi Caramaschi, 2010919. Elachistocleis carvalhoi Caramaschi, 2010920. Elachistocleis cesarii (Miranda Ribeiro (1920)921. Elachistocleis helianneae Caramaschi, 2010922. Elachistocleis erythrogaster Kwet & Di-Bernardo, 1998923. Elachistocleis magna Toledo, 2010924. Elachistocleis matogrosso Caramaschi, 2010925. Elachistocleis muiraquitan Nunes-de-Almeida & Toledo,

2012926. Elachistocleis ovalis (Schneider, 1799)927. Elachistocleis piauiensis Caramaschi & Jim, 1983

928. Elachistocleis surumu Caramaschi, 2010929. Hamptophryne alios (Wild, 1995)930. Hamptophryne boliviana (Parker, 1927)931. Myersiella microps (Dumril & Bibron, 1841)932. Stereocyclops histrio (Carvalho, 1954)933. Stereocyclops incrassatus Cope, 18701869934. Stereocyclops palmipes Caramaschi, Salles & Cruz, 2012935. Stereocyclops parkeri (Wettstein, 1934)

Family Microhylidae (Otophryninae)

936. Otophryne pyburni Campbell & Clarke, 1998937. Synapturanus mirandaribeiroi Nelson & Lescure, 1975938. Synapturanus salseri Pyburn, 1975

Family Odontophrynidae

939. Macrogenioglottus alipioi Carvalho, 1946940. Odontophrynus americanus (Dumril & Bibron, 1841)941. Odontophrynus carvalhoi Savage & Cei, 1965942. Odontophrynus cultripes Reinhardt & Ltken,

18611862943. Odontophrynus lavillai Cei, 1985944. Odontophrynus maisuma Rosset, 2008945. Odontophrynus monachus Caramaschi & Napoli, 2012946. Odontophrynus salvatori Caramaschi, 1996947. Proceratophrys appendiculata (Gnther, 1873)948. Proceratophrys aridus Cruz, Nunes & Junc, 2012949. Proceratophrys avelinoi Mercadal del Barrio & Barrio,

1993950. Proceratophrys bagnoi Brando, Caramaschi, Vaz-Silva &

Campos951. Proceratophrys bigibbosa (Peters, 1872)952. Proceratophrys belzebul Dias, Amaro, Carvalho-E-Silva &

Rodrigues, 2013953. Proceratophrys boiei (Wied-Neuwied, 1825)954. Proceratophrys branti Brando, Caramaschi, Vaz-Silva &

Campos955. Proceratophrys brauni Kwet & Faivovich, 2001956. Proceratophrys caramaschii Cruz, Nunes & Junc, 2012957. Proceratophrys carranca Godinho, Moura, Lacerda &

Feio, 2013958. Proceratophrys concavitympanum Giaretta, Bernarde &

Kokubum, 2000959. Proceratophrys cristiceps (Mller, 1884 1883)960. Proceratophrys cururu Eterovick & Sazima, 1998961. Proceratophrys dibernardoi Brando, Caramaschi, Vaz-

-Silva & Campos962. Proceratophrys goyana (Miranda-Ribeiro, 1937)963. Proceratophrys huntingtoni Avila, Pansonato &, Struss-

mann, 2012964. Proceratophrys izecksohni Dias, Amaro, Carvalho-E-Silva

& Rodrigues, 2013965. Proceratophrys laticeps Izecksohn & Peixoto, 1981966. Proceratophrys melanopogon (Miranda-Ribeiro, 1926)



967. Proceratophrys minuta Napoli, Cruz, Abreu & Del-Gran-de, 2011

968. Proceratophrys moehringi Weygoldt & Peixoto, 1985969. Proceratophrys moratoi (Jim & Caramaschi 1980)970. Proceratophrys palustris Giaretta & Sazima, 1993971. Proceratophrys paviotii Cruz, Prado & Izecksohn, 2005972. Proceratophrys phyllostomus Izecksohn, Cruz & Peixoto,

1999 1998973. Proceratophrys redacta Teixeira, Amaro, Recoder, Vechio

& Rodrigues, 2012974. Proceratophrys renalis (Miranda-Ribeiro, 1920)975. Proceratophrys rotundipalpebra Martins & Giaretta, 2013976. Proceratophrys rondonae Prado & Pombal, 2008977. Proceratophrys sanctaritae Cruz & Napoli, 2010978. Proceratophrys schirchi (Miranda-Ribeiro, 1937)979. Proceratophrys strussmannae vila, Kawashita-Ribeiro &

Morais, 2011980. Proceratophrys subguttata Izecksohn, Cruz & Peixoto,

1999 1998981. Proceratophrys tupinamba Prado & Pombal, 2008982. Proceratophrys vielliardi Martins & Giaretta, 2011

Family Pipidae

983. Pipa arrabali Izecksohn, 1976984. Pipa carvalhoi (Miranda-Ribeiro, 1937)985. Pipa pipa (Linnaeus, 1758)986. Pipa snethlageae Mller, 1914

Family Ranidae

987. Lithobates catesbeianus (Shaw, 1802) INVASIVE SPECIES988. Lithobates palmipes (Spix, 1824)

ORDER CAUDATA

Family Plethodontidae (Plethodontinae)

989. Bolitoglossa altamazonica (Cope, 1874)990. Bolitoglossa caldwellae Brcko, Hoogmoed & Neckel-Oli-

veira, 2013991. Bolitoglossa madeira Brcko, Hoogmoed & Neckel-Olivei-

ra, 2013992. Bolitoglossa paraensis (Unterstein, 1930)993. Bolitoglossa tapajnica Brcko, Hoogmoed & Neckel-Oli-

veira, 2013

ORDER GYMNOPHIONA

Family Caeciliidae

994. Caecilia armata Dunn, 1942995. Caecilia gracilis Shaw, 1802

996. Caecilia mertensi Taylor, 1973997. Caecilia tentaculata Linnaeus, 1758998. Oscaecilia hypereumeces Taylor, 1968

Family Rhinatrematidae

999. Rhinatrema bivittatum (Cuvier in Gurin-Mneville, 1829)

1000. Rhinatrema ron Wilkinson & Gower, 2010

Family Siphonopidae

1001. Brasilotyphlus braziliensis (Dunn, 1945)1002. Brasilotyphlus guarantanus Maciel, Mott & Hoogmoed,

20091003. Luetkenotyphlus brasiliensis (Ltken, 1852 1851)1004. Microcaecilia rochai Maciel & Hoogmoed, 20111005. Microcaecilia marvaleewakeae Maciel & Hoogmoed,

20131006. Microcaecilia supernumeraria Taylor, 19691007. Microcaecilia trombetas Maciel & Hoogmoed, 20111008. Microcaecilia unicolor (Dumril, 1863)1009. Mimosiphonops reinhardti Wilkinson & Nussbaum,

19921010. Mimosiphonops vermiculatus Taylor, 19681011. Siphonops annulatus (Mikan, 1820)1012. Siphonops hardyi Boulenger, 18881013. Siphonops insulanus Ihering, 19111014. Siphonops leucoderus Taylor, 19681015. Siphonops paulensis Boettger, 1892

Family Typhlonectidae

1016. Atretochoana eiselti (Taylor, 1968)1017. Chthonerpeton arii Cascon & Lima-Verde, 19941018. Chthonerpeton braestrupi Taylor, 19681019. Chthonerpeton exile Nussbaum & Wilkinson, 19871020. Chthonerpeton indistinctum (Reinhardt & Ltken,

18621861)1021. Chthonerpeton noctinectes da Silva, Britto-Pereira &

Caramaschi, 20031022. Chthonerpeton perissodus Nussbaum & Wilkinson 19871023. Chthonerpeton viviparum Parker & Wettstein, 19291024. Nectocaecilia petersii (Boulenger, 1882)1025. Potomotyphlus kaupii (Berthold, 1859)1026. Typhlonectes compressicauda (Dumril & Bibron, 1841)

REFERENCES

Frost, D.R. 2014. Amphibian Species of the World: An Online Reference. Version 6.0. Electronic Database accessible at: http://research.amnh.org/herpetology/amphibia/index.html. American Museum of Natural History, New York, USA.



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