TESIS JOSE GUTIERREZ PARTE 2.pdf

AN

EXO

S

Anexo 1: Datos meteorolgicos de las islas

Canarias

321

Parmetros e ndices bioclimticos Sigla Valor

Amplitud trmica anual Ic 6,2

ndice de termicidad It 567

ndice de termicidad compensado Itc 549

ndice ombrotrmico anual Io 0,69

Temperatura positiva Tp 2508

Precipitacin positiva Pp 174

Termopluvimetra recogida por la estacin meteorolgica de Punta Orchilla (El Hierro).

Valores bioclimatolgicos de la estacin de Punta Orchilla (El Hierro).

Diagrama bioclimtico de Punta Orchilla

(El Hierro).

Anexo 1. Datos de la estacin meteorolgica de Punta Orchilla (El Hierro) (informacin extrada de Worldwide

Bioclimatic Classification System, 1996-2009, S.Rivas-Martinez & S.Rivas-Saenz, Phytosociological Research

Center, Spain. http://www.globalbioclimatics.org).

322








Termopluvimetra recogida por la estacin meteorolgica de Vallehermoso (La Gomera).

Valores bioclimatolgicos de la estacin de Vallehermoso (La Gomera).

Diagrama bioclimtico de Vallehermoso

(La Gomera).

Anexo 1. Datos de la estacin meteorolgica de Vallehermoso (La Gomera) (informacin extrada de Worldwide



323








Termopluvimetra recogida por la estacin meteorolgica de los Llanos de Aridane (La Palma).

Valores bioclimatolgicos de la estacin de Los Llanos de Aridane (La Palma).

Diagrama bioclimtico de Los Llanos de

Aridane (La Palma).

Anexo 1. Datos de la estacin meteorolgica de Los Llanos de Aridane (La Palma) (informacin extrada de

Worldwide Bioclimatic Classification System, 1996-2009, S.Rivas-Martinez & S.Rivas-Saenz, Phytosociological

Research Center, Spain. http://www.globalbioclimatics.org).

324








Valores bioclimatolgicos de la estacin de Santa Cruz (Tenerife).

Termopluvimetra recogida por la estacin meteorolgica de Santa Cruz (Tenerife).

Diagrama bioclimtico de Santa Cruz

(Tenerife).

Anexo 1. Datos de la estacin meteorolgica de Santa Cruz de Tenerife (Tenerife) (informacin extrada de



325








Diagrama bioclimtico de Izana (El Teide,

Tenerife)

Termopluvimetra recogida por la estacin meteorolgica de Izana (El Teide, Tenerife).

Valores bioclimatolgicos de la estacin de Izana (El Teide, Tenerife).

Anexo 1. Datos de la estacin meteorolgica de Izana (El Teide, Tenerife) (informacin extrada de Worldwide



326








Diagrama bioclimtico de Las Palmas de

Gran Canaria (Gran Canaria).

Termopluvimetra recogida por la estacin meteorolgica de Las Palmas de Gran Canaria (Gran Canaria).

Valores bioclimatolgicos de la estacin de Las Palmas de Gran Canaria (Gran Canaria).

Anexo 1. Datos de la estacin meteorolgica de Las Palmas de Gran Canaria (Gran Canaria) (informacin extrada

de Worldwide Bioclimatic Classification System, 1996-2009, S.Rivas-Martinez & S.Rivas-Saenz, Phytosociological


327








Diagrama bioclimtico del aeropuerto de

Fuerteventura


Valores bioclimatolgicos de la estacin del aeropuerto de Fuerteventura.

Anexo 1. Datos de la estacin meteorolgica del aeropuerto de Fuerteventura (informacin extrada de



328








Diagrama bioclimtico de Tas

(Lanzarote)


Valores bioclimatolgicos de la estacin de Tas (Lanzarote).

Anexo 1. Datos de la estacin meteorolgica de Tas (Lanzarote) (informacin extrada de Worldwide Bioclimatic

Classification System, 1996-2009, S.Rivas-Martinez & S.Rivas-Saenz, Phytosociological Research Center, Spain.

http://www.globalbioclimatics.org).

Anexo 2: Actualizacin del catlogo de

colepteros acuticos de las islas

Canarias.

331

COLEOPTERA H P G T C F L

DRYOPIDAE Dryops gracilis Karsch, 1881 H P G T C

DYTISCIDAE

Agabus biguttatus (Olivier, 1795) P G T C Agabus conspersus (Marsham, 1802) T C Agabus nebulosus (Forster, 1771) H P G T C Bidessus minutissimus (Germar, 1824) P G T C F Cybister tripunctatus africanus Laporte, 1835 T C Eretes sticticus (Linnaeus, 1767) H P T C F L Graptodytes delectus (Wollaston, 1864) T C Herophydrus musicus (Klug, 1834) H G T C F L Hydroglyphus geminus (Fabricius, 1792) F? Hydroporus compunctus Wollaton, 1865 P G T Hydroporus errans Sharp, 1882 H P G T C Hydroporus lucasi Reiche, 1866 H P G T C Hydroporus pilosus (Guignot, 1949) T C Hygrotus confluens (Fabricius, 1787) H P G T C F L Hyphydrus maculatus Babington, 1841 C Laccophilus hyalinus testaceus Aub, 1837 G T C Meladema coriacea Laporte, 1834 G T C F Meladema imbricata (Wollaston, 1871) P G T Nebrioporus canariensis (Bedel, 1881) H P G T C Nebrioporus ceresyi (Aub, 1838) F L Nebrioporus clarkii Wollaston, 1862 F Stictonectes canariensis Machado, 1987 C

GYRINIDAE

Aulonogyrus striatus (Fabricius, 1792) G T C Gyrinus dejeani Brull, 1832 G T C F Gyrinus urinator Illiger, 1807 P G T C

HALIPLIDAE

Haliplus lineatocollis Marsham, 1802 G T C HELOPHORIDAE

Helophorus longitarsis Wollaston, 1864 F L HYDRAENIDAE

Hydraena quadricollis Wollaston, 1864 T? Hydraena serricollis Wollaston, 1864 P G T C Limnebius canariensis Orchymont, 1938 C Limnebius gracilipes Wollaston, 1864 P G T C Limnebius punctatus Wollaston, 1864 T Limnebius similis Wollaston, 1865 G Ochthebius bifoveolatus Walt, 1835 F L Ochthebius heeri (Wollaston, 1864) H P G T C L Ochthebius lapidicola Wollaston, 1864 H P G T C F L Ochthebius quadrifoveolatus Wollaston, 1854 P G T C F Ochthebius rugulosus Wollaston, 1854 H P G T C F L

332

COLEOPTERA H P G T C F L

Ochthebius salinator Peyerimhoff, 1924 L Ochthebius subpictus subpictus Wollaston, 1857 F Ochthebius tivelunus Ferro, 1984 F

HYDROCHIDAE

Hydrochus grandicollis Kiesenwetter, 1870 T C HYDROPHILIDAE

Anacaena haemorrhoa (Wollaston, 1864) P G T C Berosus hispanicus Kster, 1847 F Chaetarthria similis Wollaston, 1864 P G T C F Coelostoma hispanicum (Kster, 1848) P G T C F L Enochrus bicolor (Fabricius, 1792) P G T C F L Enochrus politus (Kster, 1849) G T C F L Helochares lividus (Forster, 1771) H G T C Laccobius atrocephalus canariensis Orchymont, 1940

P G T C F L

Laccobius praecipuus Kuwert, 1890 P T C Paracymus relaxus Rey, 1884 F L

Anexo 3: Matriz de abundancia de especies de

colepteros acuticos de las islas

Canarias

335

ESTACIN 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24

Hal. lin. 9 1 11 11 40 1

Aul. str.

Gyr. dej. 3 26 38 16 4

Gyr. uri. 11 8

Aga. big. 4 1 1 1 8 2

Aga. con.

Aga. neb. 3 1 1 3 3

Mel. cor. 2 23 6 12 1

Mel. imb.

Cyb. tri.

Ere. sti.

Bid. min. 55 2 1 27 4

Hyd. gem.

Gra. del. 36 86 36 3

Hyd. com.

Hyd. err. 9 2 2 34 57

Hyd. luc.

Hyd. pil.

Neb. cla. 2 1 24 15 1

Neb. can. 3

Neb. cer. 48 20 27 39 1 2 24 19

Sti. can. 67 12

Her. mus. 4 7 22 14 7

Hyg. con. 4 2 10 2 6 19 1

Hyp. mac. 19 10

Lac. hyl. 6 4 65 13 29

Ana. hae. 1 1 1

Par. rel. 3 6 1 1 3

Ber. his. 12 12 22

Cha. sim.

Eno. bic. 2 4 6 3

Eno. pol. 1 5 6 33 10 20 8 29 41 4 7 11

Hel. liv. 2 3 21 17 15

Lac. atr. 28 4 13 2 11 2 5 2 8 1 38

Lac. pra. 1

Coe. his. 9 1 1 2 40

Hel. lon. 3 1 1

Hyd. gra. 79 2

Hyd. ser. 20

Hyd. qua.

Lim. can. 1 5 6 5

Lim. gra. 39

Lim. pun.

Lim. sim.

Och. rug. 2 40 5 3 17 13 18

Och. hee.

Och. bif. 18 1 23

Och. lap.

Och. qua. 9 4 36 3

Och. sal. 6

Och. sub. 10 7 3 28 26

Och. tiv. 3

Dry. gra. 3 18 58 39 15 15 17

336

ESTACIN 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49

Hal. lin. 3 1 2

Aul. str.

Gyr. dej. 34 23 15 5 12 1 11 16 22 6

Gyr. uri. 2 3 10 13 15 2 7 21 2

Aga. big. 6 2 64 9 12 2 4 13 4 6

Aga. con.

Aga. neb. 58 6 2 14 24 1 6 5 3

Mel. cor. 3 4 1 2 2

Mel. imb.

Cyb. tri. 3

Ere. sti.

Bid. min. 1 7

Hyd. gem.

Gra. del. 14 34 9 9

Hyd. com.

Hyd. err. 9 38 6 20 15 5 8 7 15 4 6

Hyd. luc. 5 2 2

Hyd. pil.

Neb. cla.

Neb. can. 24 13 10 2 5 3 12

Neb. cer.

Sti. can. 1

Her. mus. 18 4 4

Hyg. con. 1 1 5 12

Hyp. mac.

Lac. hyl. 2 10 2 1

Ana. hae.

Par. rel.

Ber. his.

Cha. sim. 3

Eno. bic. 1 4

Eno. pol. 4 5

Hel. liv. 7 2

Lac. atr. 1 7 2 23 5 1 13 4 5 2

Lac. pra. 1 7

Coe. his. 1 5

Hel. lon.

Hyd. gra. 10

Hyd. ser.

Hyd. qua.

Lim. can. 1

Lim. gra. 4 1 4

Lim. pun.

Lim. sim.

Och. rug. 2 30 30 16 1 2 3 1 15 13 3 10

Och. hee.

Och. bif.

Och. lap. 1 1

Och. qua. 2 17 1 1 7 9

Och. sal.

Och. sub.

Och. tiv.

Dry. gra. 21 2 1 1 2 1 4 4 1

337

ESTACIN 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71

Hal. lin.

Aul. str.

Gyr. dej. 19

Gyr. uri. 7

Aga. big. 4 1

Aga. con.

Aga. neb. 13 8

Mel. cor.

Mel. imb.

Cyb. tri.

Ere. sti.

Bid. min. 2 3

Hyd. gem.

Gra. del.

Hyd. com.

Hyd. err. 3 53

Hyd. luc. 4

Hyd. pil.

Neb. cla.

Neb. can. 1 2

Neb. cer.

Sti. can.

Her. mus. 36

Hyg. con. 9 1

Hyp. mac.

Lac. hyl. 7

Ana. hae. 1

Par. rel.

Ber. his.

Cha. sim.

Eno. bic. 17

Eno. pol. 4

Hel. liv. 1 2

Lac. atr. 7 18 30 16 2 1

Lac. pra. 1 5 4

Coe. his. 5 10

Hel. lon.

Hyd. gra.

Hyd. ser.

Hyd. qua.

Lim. can.

Lim. gra. 1

Lim. pun.

Lim. sim.

Och. rug. 1 11 3 4

Och. hee. 18

Och. bif.

Och. lap. 1 14

Och. qua. 1 2 30 12

Och. sal.

Och. sub.

Och. tiv.

Dry. gra. 2 1

Anexo 4: Mapas de distribucin de las especies

estudiadas.

!(

!(!(

!(

!(

!(

!(

!(!(

!(

!( !(

!(

!(

250000,000000

250000,000000

270000,000000

270000,000000

290000,000000

290000,000000

310000,000000

310000,000000

330000,000000

330000,000000

350000,000000

350000,000000

370000,000000

370000,000000

390000,000000

390000,000000

410000,000000

410000,000000

430000,000000

430000,000000

450000,000000

450000,000000

470000,000000

470000,000000

490000,000000

490000,000000

510000,000000

510000,000000

306000

0,00000

0

306000

0,00000

0

308000

0,00000

0

308000

0,00000

0

310000

0,00000

0

310000

0,00000

0

312000

0,00000

0

312000

0,00000

0

314000

0,00000

0

314000

0,00000

0

316000

0,00000

0

316000

0,00000

0

318000

0,00000

0

318000

0,00000

0

320000

0,00000

0

320000

0,00000

0

322000

0,00000

0

322000

0,00000

0

!( Haliplus lineatocollisIslas Canarias

!(!(

!(!(

!(

!(!(!(

!(!(

!(!( !(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(!(

!(

!(

260000,000000

260000,000000

280000,000000

280000,000000

300000,000000

300000,000000

320000,000000

320000,000000

340000,000000

340000,000000

360000,000000

360000,000000

380000,000000

380000,000000

400000,000000

400000,000000

420000,000000

420000,000000

440000,000000

440000,000000

460000,000000

460000,000000

480000,000000

480000,000000

500000,000000

500000,000000

520000,000000

520000,000000

540000,000000

540000,000000

560000,000000

560000,000000

580000,000000

580000,000000

600000,000000

600000,000000

620000,000000

620000,000000 301000

0,00000

0303000

0,00000

0

304000

0,00000

0306000

0,00000

0

307000

0,00000

0309000

0,00000

0

310000

0,00000

0312000

0,00000

0

313000

0,00000

0315000

0,00000

0

316000

0,00000

0318000

0,00000

0

319000

0,00000

0321000

0,00000

0

322000

0,00000

0324000

0,00000

0

325000

0,00000

0

Islas Canarias!( Gyrinus dejeani

60 0 6030 km

190000,000000

190000,000000

210000,000000

210000,000000

230000,000000

230000,000000

250000,000000

250000,000000

270000,000000

270000,000000

290000,000000

290000,000000

310000,000000

310000,000000

330000,000000

330000,000000

350000,000000

350000,000000

370000,000000

370000,000000

390000,000000

390000,000000

410000,000000

410000,000000

430000,000000

430000,000000

450000,000000

450000,000000

470000,000000

470000,000000

490000,000000

490000,000000

510000,000000

510000,000000

530000,000000

530000,000000 304000

0,00000

0

305000

0,00000

0

306000

0,00000

0

307000

0,00000

0

308000

0,00000

0

309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

324000

0,00000

0

325000

0,00000

0

326000

0,00000

0

327000

0,00000

0

328000

0,00000

0

Islas Canarias

Gyrinus urinator

50 0 5025 km

!(!(!(

!(

!(

!(!(!(!(

!(

!(!(!(

!(

!( !(

!(

!(

!(!(

!(

!(!(

!(

!(

!(

!(

!(!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

190000,000000

190000,000000

210000,000000

210000,000000

230000,000000

230000,000000

250000,000000

250000,000000

270000,000000

270000,000000

290000,000000

290000,000000

310000,000000

310000,000000

330000,000000

330000,000000

350000,000000

350000,000000

370000,000000

370000,000000

390000,000000

390000,000000

410000,000000

410000,000000

430000,000000

430000,000000

450000,000000

450000,000000

470000,000000

470000,000000

490000,000000

490000,000000

510000,000000

510000,000000

530000,000000

530000,000000 304000

0,00000

0

305000

0,00000

0

306000

0,00000

0

307000

0,00000

0

308000

0,00000

0

309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

324000

0,00000

0

325000

0,00000

0

326000

0,00000

0

327000

0,00000

0

328000

0,00000

0

Islas Canarias

!( Agabus biguttatus

50 0 5025 km

!(!(!(

!(

!(

!(!(!(!(

!(

!(!(!(

!(

!( !(

!(

!(

!(!(

!(

!(!(

!(

!(

!(

!(

!(!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(!(

!(

!(

!(

!(

!(!(

!(

!(

!(!(

!(

!(

!(!(

!(

!(

!(!(

!(

!(

!(

!(

!(

!(

!(

!(!(!(

!(!(!(

!(

190000,000000

190000,000000

210000,000000

210000,000000

230000,000000

230000,000000

250000,000000

250000,000000

270000,000000

270000,000000

290000,000000

290000,000000

310000,000000

310000,000000

330000,000000

330000,000000

350000,000000

350000,000000

370000,000000

370000,000000

390000,000000

390000,000000

410000,000000

410000,000000

430000,000000

430000,000000

450000,000000

450000,000000

470000,000000

470000,000000

490000,000000

490000,000000

510000,000000

510000,000000

530000,000000

530000,000000 304000

0,00000

0

305000

0,00000

0

306000

0,00000

0

307000

0,00000

0

308000

0,00000

0

309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

324000

0,00000

0

325000

0,00000

0

326000

0,00000

0

327000

0,00000

0

328000

0,00000

0

Islas Canarias

!( Agabus nebulosus

50 0 5025 km

!(!(

!(!(

!(!(

!( !(!(

!(

!(

!(

!(!(

!(

!(

250000,000000

250000,000000

270000,000000

270000,000000

290000,000000

290000,000000

310000,000000

310000,000000

330000,000000

330000,000000

350000,000000

350000,000000

370000,000000

370000,000000

390000,000000

390000,000000

410000,000000

410000,000000

430000,000000

430000,000000

450000,000000

450000,000000

470000,000000

470000,000000

490000,000000

490000,000000

510000,000000

510000,000000

530000,000000

530000,000000

550000,000000

550000,000000

570000,000000

570000,000000

590000,000000

590000,000000

610000,000000

610000,000000 30200

00,0000

00

303000

0,00000

0 305000

0,00000

0

306000

0,00000

0 308000

0,00000

0

309000

0,00000

0 311000

0,00000

0

312000

0,00000

0 314000

0,00000

0

315000

0,00000

0 317000

0,00000

0

318000

0,00000

0 320000

0,00000

0

321000

0,00000

0 323000

0,00000

0

324000

0,00000

0 326000

0,00000

0

327000

0,00000

0

Islas Canarias !( Meladema coriacea

40 0 4020 km

!(

!(

!(

!(

170000,000000

170000,000000

190000,000000

190000,000000

210000,000000

210000,000000

230000,000000

230000,000000

250000,000000

250000,000000

270000,000000

270000,000000

290000,000000

290000,000000

310000,000000

310000,000000

330000,000000

330000,000000

350000,000000

350000,000000

370000,000000

370000,000000

390000,000000

390000,000000

410000,000000

410000,000000 308000

0,00000

0

309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

324000

0,00000

0

325000

0,00000

0

Islas Canarias

!( Meladema imbricata

30 0 3015 km

!(!(

!(

310000,000000

310000,000000

320000,000000

320000,000000

330000,000000

330000,000000

340000,000000

340000,000000

350000,000000

350000,000000

360000,000000

360000,000000

370000,000000

370000,000000

380000,000000

380000,000000

390000,000000

390000,000000

400000,000000

400000,000000

410000,000000

410000,000000

420000,000000

420000,000000

430000,000000

430000,000000

440000,000000

440000,000000

450000,000000

450000,000000309000

0,00000

0

309000

0,00000

031000

00,0000

00

310000

0,00000

031100

00,0000

00

311000

0,00000

031200

00,0000

00

312000

0,00000

031300

00,0000

00

313000

0,00000

031400

00,0000

00

314000

0,00000

031500

00,0000

00

315000

0,00000

031600

00,0000

00

316000

0,00000

031700

00,0000

00

317000

0,00000

031800

00,0000

00

318000

0,00000

031900

00,0000

00

319000

0,00000

0

Islas Canarias

!( Cybister tripunctatus africanus

6 0 63 km

!(

!(

200000,000000

200000,000000

210000,000000

210000,000000

220000,000000

220000,000000

230000,000000

230000,000000

240000,000000

240000,000000

250000,000000

250000,000000

260000,000000

260000,000000

270000,000000

270000,000000

315000

0,00000

0

315000

0,00000

0

316000

0,00000

0

316000

0,00000

0

317000

0,00000

0

317000

0,00000

0

318000

0,00000

0

318000

0,00000

0

319000

0,00000

0

319000

0,00000

0

Islas Canarias

!( Eretes sticticus

4 0 42 km

!(

!(!(

!(!(

!(!(

!(

!(!(

!(!(

!(

!(

200000,000000

200000,000000

230000,000000

230000,000000

260000,000000

260000,000000

290000,000000

290000,000000

320000,000000

320000,000000

350000,000000

350000,000000

380000,000000

380000,000000

410000,000000

410000,000000

440000,000000

440000,000000

470000,000000

470000,000000

500000,000000

500000,000000

530000,000000

530000,000000

560000,000000

560000,000000

590000,000000

590000,000000

620000,000000

620000,00000030200

00,0000

00

303000

0,00000

0 305000

0,00000

0

306000

0,00000

0 308000

0,00000

0

309000

0,00000

0 311000

0,00000

0

312000

0,00000

0 314000

0,00000

0

315000

0,00000

0 317000

0,00000

0

318000

0,00000

0 320000

0,00000

0

321000

0,00000

0 323000

0,00000

0

324000

0,00000

0 326000

0,00000

0

327000

0,00000

0 329000

0,00000

0

330000

0,00000

0

Islas Canarias

!( Bidessus minutissimus

20 0 2010 km

!(!(

!(

!(

!(

!(

!(

!(

!(

!(

310000,000000

310000,000000

320000,000000

320000,000000

330000,000000

330000,000000

340000,000000

340000,000000

350000,000000

350000,000000

360000,000000

360000,000000

370000,000000

370000,000000

380000,000000

380000,000000

390000,000000

390000,000000

400000,000000

400000,000000

410000,000000

410000,000000

420000,000000

420000,000000

430000,000000

430000,000000

440000,000000

440000,000000

450000,000000

450000,000000

460000,000000

460000,000000

470000,000000

470000,000000

480000,000000

480000,000000

490000,000000

490000,000000

306000

0,00000

0

306000

0,00000

0

308000

0,00000

0

308000

0,00000

0

310000

0,00000

0

310000

0,00000

0

312000

0,00000

0

312000

0,00000

0

314000

0,00000

0

314000

0,00000

0

316000

0,00000

0

316000

0,00000

0

318000

0,00000

0

318000

0,00000

0

Islas Canarias

!( Graptodytes delectus

20 0 2010 km

!(

200000,000000

200000,000000

210000,000000

210000,000000

220000,000000

220000,000000

230000,000000

230000,000000

240000,000000

240000,000000

250000,000000

250000,000000

260000,000000

260000,000000

270000,000000

270000,000000

315000

0,00000

0

315000

0,00000

0

316000

0,00000

0

316000

0,00000

0

317000

0,00000

0

317000

0,00000

0

318000

0,00000

0

318000

0,00000

0

319000

0,00000

0

319000

0,00000

0

Islas Canarias

!( Hydroporus compunctus

4 0 42 km

!(

!(!(!(!(

!(

!(

!(

!(

!(

!(

!(!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

!(

180000,000000

180000,000000

200000,000000

200000,000000

220000,000000

220000,000000

240000,000000

240000,000000

260000,000000

260000,000000

280000,000000

280000,000000

300000,000000

300000,000000

320000,000000

320000,000000

340000,000000

340000,000000

360000,000000

360000,000000

380000,000000

380000,000000

400000,000000

400000,000000

420000,000000

420000,000000

440000,000000

440000,000000

460000,000000

460000,000000

480000,000000

480000,000000

500000,000000

500000,000000

520000,000000

520000,000000 3010

000,00

0000

302000

0,00000

0

303000

0,00000

0

304000

0,00000

0

305000

0,00000

0

306000

0,00000

0

307000

0,00000

0

308000

0,00000

0

309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

324000

0,00000

0

Islas Canarias

!( Hydroporus errans

50 0 5025 km

!(

!(

!(!(

180000,000000

180000,000000

200000,000000

200000,000000

220000,000000

220000,000000

240000,000000

240000,000000

260000,000000

260000,000000

280000,000000

280000,000000

300000,000000

300000,000000

320000,000000

320000,000000

340000,000000

340000,000000

360000,000000

360000,000000

380000,000000

380000,000000 305000

0,00000

0306000

0,00000

0 307000

0,00000

0308000

0,00000

0 309000

0,00000

0310000

0,00000

0 311000

0,00000

0312000

0,00000

0 313000

0,00000

0314000

0,00000

0 315000

0,00000

0316000

0,00000

0 317000

0,00000

0318000

0,00000

0 319000

0,00000

0

Islas Canarias

!( Hydroporus lucasi

7 0 73,5 km

!(

!(

310000,000000

310000,000000

320000,000000

320000,000000

330000,000000

330000,000000

340000,000000

340000,000000

350000,000000

350000,000000

360000,000000

360000,000000

370000,000000

370000,000000

380000,000000

380000,000000

390000,000000

390000,000000

400000,000000

400000,000000

410000,000000

410000,000000

420000,000000

420000,000000

430000,000000

430000,000000

440000,000000

440000,000000

450000,000000

450000,000000

460000,000000

460000,000000

470000,000000

470000,000000

480000,000000

480000,000000

490000,000000

490000,000000

306000

0,00000

0

306000

0,00000

0

308000

0,00000

0

308000

0,00000

0

310000

0,00000

0

310000

0,00000

0

312000

0,00000

0

312000

0,00000

0

314000

0,00000

0

314000

0,00000

0

316000

0,00000

0

316000

0,00000

0

318000

0,00000

0

318000

0,00000

0

Islas Canarias

!( Hydroporus pilosus

20 0 2010 km

!(

!(

!(!(

!(

530000,000000

530000,000000

540000,000000

540000,000000

550000,000000

550000,000000

560000,000000

560000,000000

570000,000000

570000,000000

580000,000000

580000,000000

590000,000000

590000,000000

600000,000000

600000,000000

610000,000000

610000,000000

620000,000000

620000,000000

630000,000000

630000,000000

640000,000000

640000,000000

650000,000000

650000,000000

660000,000000

660000,000000

670000,000000

670000,000000

680000,000000

680000,000000

309000

0,00000

0

309000

0,00000

0310000

0,00000

0

310000

0,00000

0311000

0,00000

0

311000

0,00000

0312000

0,00000

0

312000

0,00000

0313000

0,00000

0

313000

0,00000

0314000

0,00000

0

314000

0,00000

0315000

0,00000

0

315000

0,00000

0316000

0,00000

0

316000

0,00000

0317000

0,00000

0

317000

0,00000

0318000

0,00000

0

318000

0,00000

0319000

0,00000

0

319000

0,00000

0320000

0,00000

0

320000

0,00000

0

Islas Canarias

!( Nebrioporus clarkii

8 0 84 km

!(!(

!(!(

!(

!(

!(

!(!(

!(!(

!(!(

!(!(

!(!(

!(

!(!(

!(

!(!(

!(!(

!(

!(

!(

210000,000000

210000,000000

230000,000000

230000,000000

250000,000000

250000,000000

270000,000000

270000,000000

290000,000000

290000,000000

310000,000000

310000,000000

330000,000000

330000,000000

350000,000000

350000,000000

370000,000000

370000,000000

390000,000000

390000,000000

410000,000000

410000,000000

430000,000000

430000,000000

450000,000000

450000,000000304000

0,00000

0

304000

0,00000

0

306000

0,00000

0

306000

0,00000

0

308000

0,00000

0

308000

0,00000

0

310000

0,00000

0

310000

0,00000

0

312000

0,00000

0

312000

0,00000

0

314000

0,00000

0

314000

0,00000

0

316000

0,00000

0

316000

0,00000

0

318000

0,00000

0

318000

0,00000

0

320000

0,00000

0

320000

0,00000

0

322000

0,00000

0

322000

0,00000

0

Islas Canarias!( Nebrioporus canariensis

10 0 105 km

!(

!(

!(

!(!(

!(

!(

!(

!(

!(

540000,000000

540000,000000

560000,000000

560000,000000

580000,000000

580000,000000

600000,000000

600000,000000

620000,000000

620000,000000

640000,000000

640000,000000

660000,000000

660000,000000

680000,000000

680000,000000

700000,000000

700000,000000

720000,000000

720000,000000

740000,000000

740000,000000 3100

000,00

0000311000

0,00000

0 312000

0,00000

0313000

0,00000

0

314000

0,00000

0315000

0,00000

0

316000

0,00000

0317000

0,00000

0 318000

0,00000

0

319000

0,00000

0 320000

0,00000

0

321000

0,00000

0 322000

0,00000

0323000

0,00000

0

324000

0,00000

0

Islas Canarias

!( Nebrioporus ceresyi

10 0 105 km

!(

!(

!(

!(!(

420000,000000

420000,000000

430000,000000

430000,000000

440000,000000

440000,000000

450000,000000

450000,000000

460000,000000

460000,000000

470000,000000

470000,000000

480000,000000

480000,000000

490000,000000

490000,000000

500000,000000

500000,000000

307000

0,00000

0

307000

0,00000

0

308000

0,00000

0

308000

0,00000

0

309000

0,00000

0

309000

0,00000

0

310000

0,00000

0

310000

0,00000

0

311000

0,00000

0

311000

0,00000

0

312000

0,00000

0

312000

0,00000

0

Islas Canarias

!( Stictonectes canariensis

10 0 105 km

!(

!(

!(

!(

!(!(

!(

!(

!(

!(

!(

!(

!(

180000,000000

180000,000000

210000,000000

210000,000000

240000,000000

240000,000000

270000,000000

270000,000000

300000,000000

300000,000000

330000,000000

330000,000000

360000,000000

360000,000000

390000,000000

390000,000000

420000,000000

420000,000000

450000,000000

450000,000000

480000,000000

480000,000000

510000,000000

510000,000000

540000,000000

540000,000000

570000,000000

570000,000000

600000,000000

600000,000000

630000,000000

630000,000000

660000,000000

660000,000000

299000

0,00000

0

301000

0,00000

0

302000

0,00000

0

304000

0,00000

0

305000

0,00000

0

307000

0,00000

0

308000

0,00000

0

310000

0,00000

0

311000

0,00000

0

313000

0,00000

0

314000

0,00000

0

316000

0,00000

0

317000

0,00000

0

319000

0,00000

0

320000

0,00000

0

322000

0,00000

0

323000

0,00000

0

325000

0,00000

0

326000

0,00000

0

328000

0,00000

0

329000

0,00000

0

331000

0,00000

0

332000

0,00000

0

Islas Canarias !( Herophydrus musicus

50 0 5025 km

!(!(

!(

!(!(

!(

!(

!(

!(!(

!(

!(

!(

!(!(

!(

!(

!(

180000,000000

180000,000000

210000,000000

210000,000000

240000,000000

240000,000000

270000,000000

270000,000000

300000,000000

300000,000000

330000,000000

330000,000000

360000,000000

360000,000000

390000,000000

390000,000000

420000,000000

420000,000000

450000,000000

450000,000000

480000,000000

480000,000000

510000,000000

510000,000000

540000,000000

540000,000000

570000,000000

570000,000000

600000,000000

600000,000000

630000,000000

630000,000000

660000,000000

660000,000000

299000

0,00000

0

301000

0,00000

0

302000

0,00000

0

304000

0,00000

0

305000

0,00000

0

307000

0,00000

0

308000

0,00000

0

310000

0,00000

0

311000

0,00000

0

313000

0,00000

0

314000

0,00000

0

316000

0,00000

0

317000

0,00000

0

319000

0,00000

0

320000

0,00000

0

322000

0,00000

0

323000

0,00000

0

325000

0,00000

0

326000

0,00000

0

328000

0,00000

0

329000

0,00000

0

331000

0,00000

0

332000

0,00000

0

Islas Canarias !( Hygrotus confluens

50 0 5025 km

!(!(

420000,000000

420000,000000

430000,000000

430000,000000

440000,000000

440000,000000

450000,000000

450000,000000

460000,000000

460000,000000

470000,000000

470000,000000

480000,000000

480000,000000

490000,000000

490000,000000

500000,000000

500000,000000

307000

0,00000

0

307000

0,00000

0

308000

0,00000

0

308000

0,00000

0

309000

0,00000

0

309000

0,00000

0

310000

0,00000

0

310000

0,00000

0

311000

0,00000

0

311000

0,00000

0

312000

0,00000

0

312000

0,00000

0

Islas Canarias

!( Hyphydrus maculatus

10 0 105 km

!( !(

!(!(

!(

!(

!(!(

!(

!(

!(

!(

260000,000000

260000,000000

280000,000000

280000,000000

300000,000000

300000,000000

320000,000000

320000,000000

340000,000000

340000,000000

360000,000000

360000,000000

380000,000000

380000,000000

400000,000000

400000,000000

420000,000000

420000,000000

440000,000000

440000,000000

460000,000000

460000,000000

480000,000000

480000,000000305000

0,00000

0

305000

0,00000

0

307000

0,00000

0

307000

0,00000

0

309000

0,00000

0

309000

0,00000

0

311000

0,00000

0

311000

0,00000

0

313000

0,00000

0

313000

0,00000

0

315000

0,00000

0

315000

0,00000

0

317000

0,00000

0

317000

0,00000

0

319000

0,00000

0

319000

0,00000

0

321000

0,00000

0

321000

0,00000

0

Islas Canarias

!( Laccophilus hyalinus testaceus

9 0 94,5 km

!(

!(

!(

!(!(

!(!(

190000,000000

190000,000000

210000,000000

210000,000000

230000,000000

230000,000000

250000,000000

250000,000000

270000,000000

270000,000000

290000,000000

290000,000000

310000,000000

310000,000000

330000,000000

330000,000000

350000,000000

350000,000000

370000,000000

370000,000000

390000,000000

390000,000000

410000,000000

410000,000000

430000,000000

430000,000000

450000,000000

450000,000000

470000,000000

470000,000000

490000,000000

490000,000000

510000,000000

510000,000000

530000,000000

530000,000000 304000

0,00000

0

305000

0,00000

0

306000

0,00000

0

307000

0,00000

0

308000

0,00000

0

309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

324000

0,00000

0

325000

0,00000

0

326000

0,00000

0

327000

0,00000

0

328000

0,00000

0

Islas Canarias

!( Anacaena haemorrhoa

50 0 5025 km

!(

!(

!(

!(

!(

520000,000000

520000,000000

540000,000000

540000,000000

560000,000000

560000,000000

580000,000000

580000,000000

600000,000000

600000,000000

620000,000000

620000,000000

640000,000000

640000,000000

660000,000000

660000,000000

680000,000000

680000,000000

700000,000000

700000,000000

720000,000000

720000,000000

740000,000000

740000,000000

760000,000000

760000,000000 309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

324000

0,00000

0

325000

0,00000

0

Islas Canarias

!( Paracymus relaxus

40 0 4020 km

!(

!(

!(

!(

!(

510000,000000

510000,000000

520000,000000

520000,000000

530000,000000

530000,000000

540000,000000

540000,000000

550000,000000

550000,000000

560000,000000

560000,000000

570000,000000

570000,000000

580000,000000

580000,000000

590000,000000

590000,000000

600000,000000

600000,000000

610000,000000

610000,000000

620000,000000

620000,000000

630000,000000

630000,000000

640000,000000

640000,000000

650000,000000

650000,000000

660000,000000

660000,000000

670000,000000

670000,000000

680000,000000

680000,000000

690000,000000

690000,000000

310000

0,00000

0

310000

0,00000

0

312000

0,00000

0

312000

0,00000

0

314000

0,00000

0

314000

0,00000

0

316000

0,00000

0

316000

0,00000

0

318000

0,00000

0

318000

0,00000

0

320000

0,00000

0

320000

0,00000

0

322000

0,00000

0

322000

0,00000

0

Islas Canarias

!( Berosus hispanicus

30 0 3015 km

!(!(

!(

190000,000000

190000,000000

210000,000000

210000,000000

230000,000000

230000,000000

250000,000000

250000,000000

270000,000000

270000,000000

290000,000000

290000,000000

310000,000000

310000,000000

330000,000000

330000,000000

350000,000000

350000,000000

370000,000000

370000,000000

390000,000000

390000,000000

410000,000000

410000,000000 309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

324000

0,00000

0

Islas Canarias

!( Chaetarthria similis

30 0 3015 km

!(

!(

!(!(

!(

!( !(

!( !(

200000,000000

200000,000000

230000,000000

230000,000000

260000,000000

260000,000000

290000,000000

290000,000000

320000,000000

320000,000000

350000,000000

350000,000000

380000,000000

380000,000000

410000,000000

410000,000000

440000,000000

440000,000000

470000,000000

470000,000000

500000,000000

500000,000000

530000,000000

530000,000000

560000,000000

560000,000000

590000,000000

590000,000000

620000,000000

620000,000000

650000,000000

650000,000000 299000

0,00000

0

301000

0,00000

0

302000

0,00000

0

304000

0,00000

0

305000

0,00000

0

307000

0,00000

0

308000

0,00000

0

310000

0,00000

0

311000

0,00000

0

313000

0,00000

0

314000

0,00000

0

316000

0,00000

0

317000

0,00000

0

319000

0,00000

0

320000

0,00000

0

322000

0,00000

0

323000

0,00000

0

325000

0,00000

0

326000

0,00000

0

328000

0,00000

0

329000

0,00000

0

Islas Canarias

!( Enochrus bicolor

25 0 2512,5 km

!(

!(!(

!(

!(!(

!(!(

!(

!(

!(

!(

!(

!(

!(

260000,000000

260000,000000

290000,000000

290000,000000

320000,000000

320000,000000

350000,000000

350000,000000

380000,000000

380000,000000

410000,000000

410000,000000

440000,000000

440000,000000

470000,000000

470000,000000

500000,000000

500000,000000

530000,000000

530000,000000

560000,000000

560000,000000

590000,000000

590000,000000

620000,000000

620000,000000

650000,000000

650000,00000030300

00,0000

00

303000

0,00000

0

306000

0,00000

0

306000

0,00000

0

309000

0,00000

0

309000

0,00000

0

312000

0,00000

0

312000

0,00000

0

315000

0,00000

0

315000

0,00000

0

318000

0,00000

0

318000

0,00000

0

321000

0,00000

0

321000

0,00000

0

324000

0,00000

0

324000

0,00000

0

327000

0,00000

0

327000

0,00000

0

330000

0,00000

0

330000

0,00000

0

Islas Canarias

!( Enochrus politus

30 0 3015 km

!(

!(!( !(

!(

!(

!(!(

!(

!(

!(

!(

!(

180000,000000

180000,000000

200000,000000

200000,000000

220000,000000

220000,000000

240000,000000

240000,000000

260000,000000

260000,000000

280000,000000

280000,000000

300000,000000

300000,000000

320000,000000

320000,000000

340000,000000

340000,000000

360000,000000

360000,000000

380000,000000

380000,000000

400000,000000

400000,000000

420000,000000

420000,000000

440000,000000

440000,000000

460000,000000

460000,000000

480000,000000

480000,000000304000

0,00000

0

304000

0,00000

0306000

0,00000

0

306000

0,00000

0308000

0,00000

0

308000

0,00000

0310000

0,00000

0

310000

0,00000

0312000

0,00000

0

312000

0,00000

0314000

0,00000

0

314000

0,00000

0316000

0,00000

0

316000

0,00000

0318000

0,00000

0

318000

0,00000

0320000

0,00000

0

320000

0,00000

0322000

0,00000

0

322000

0,00000

0324000

0,00000

0

324000

0,00000

0

Islas Canarias

!( Helochares lividus

20 0 2010 km

!(!(!( !(

!(!(

!(

!(!(!(

!(

!(!(

!(

!(!(

!(

!(

!(

!(!(

!(

!(

!(

!(

!(!(

!(

!(

!(

!(

!(!(!(

!(

180000,000000

180000,000000

210000,000000

210000,000000

240000,000000

240000,000000

270000,000000

270000,000000

300000,000000

300000,000000

330000,000000

330000,000000

360000,000000

360000,000000

390000,000000

390000,000000

420000,000000

420000,000000

450000,000000

450000,000000

480000,000000

480000,000000

510000,000000

510000,000000

540000,000000

540000,000000

570000,000000

570000,000000

600000,000000

600000,000000

630000,000000

630000,000000

660000,000000

660000,000000 299000

0,00000

0

300000

0,00000

0

302000

0,00000

0

303000

0,00000

0

305000

0,00000

0

306000

0,00000

0

308000

0,00000

0

309000

0,00000

0

311000

0,00000

0

312000

0,00000

0

314000

0,00000

0

315000

0,00000

0

317000

0,00000

0

318000

0,00000

0

320000

0,00000

0

321000

0,00000

0

323000

0,00000

0

324000

0,00000

0

326000

0,00000

0

327000

0,00000

0

329000

0,00000

0

330000

0,00000

0

332000

0,00000

0

Islas Canarias !( Laccobius atrocephalus canariensis

20 0 2010 km

!(!(!(

!(!(

!(

!(

!(

200000,000000

200000,000000

220000,000000

220000,000000

240000,000000

240000,000000

260000,000000

260000,000000

280000,000000

280000,000000

300000,000000

300000,000000

320000,000000

320000,000000

340000,000000

340000,000000

360000,000000

360000,000000

380000,000000

380000,000000

400000,000000

400000,000000

420000,000000

420000,000000

440000,000000

440000,000000

460000,000000

460000,000000 305000

0,00000

0

306000

0,00000

0

307000

0,00000

0

308000

0,00000

0

309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

Islas Canarias

!( Laccobius praecipuus

9 0 94,5 km

!(

!(!(!(

!(

!(!(

!(

!(!(

!(

!(

!(

!(!(

!(

!(

!(

!(!(

!(

200000,000000

200000,000000

230000,000000

230000,000000

260000,000000

260000,000000

290000,000000

290000,000000

320000,000000

320000,000000

350000,000000

350000,000000

380000,000000

380000,000000

410000,000000

410000,000000

440000,000000

440000,000000

470000,000000

470000,000000

500000,000000

500000,000000

530000,000000

530000,000000

560000,000000

560000,000000

590000,000000

590000,000000

620000,000000

620000,00000030200

00,0000

00

303000

0,00000

0 305000

0,00000

0

306000

0,00000

0 308000

0,00000

0

309000

0,00000

0 311000

0,00000

0

312000

0,00000

0 314000

0,00000

0

315000

0,00000

0 317000

0,00000

0

318000

0,00000

0 320000

0,00000

0

321000

0,00000

0 323000

0,00000

0

324000

0,00000

0 326000

0,00000

0

327000

0,00000

0 329000

0,00000

0

330000

0,00000

0

Islas Canarias

!( Coleostoma hispanicum

20 0 2010 km

!(

!(

!(

520000,000000

520000,000000

540000,000000

540000,000000

560000,000000

560000,000000

580000,000000

580000,000000

600000,000000

600000,000000

620000,000000

620000,000000

640000,000000

640000,000000

660000,000000

660000,000000

680000,000000

680000,000000

700000,000000

700000,000000

720000,000000

720000,000000

740000,000000

740000,000000

760000,000000

760000,000000 309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

324000

0,00000

0

325000

0,00000

0

Islas Canarias

!( Helophorus longitarsis

40 0 4020 km

!(

!(!(

!(

!(

!(

!(

200000,000000

200000,000000

220000,000000

220000,000000

240000,000000

240000,000000

260000,000000

260000,000000

280000,000000

280000,000000

300000,000000

300000,000000

320000,000000

320000,000000

340000,000000

340000,000000

360000,000000

360000,000000

380000,000000

380000,000000

400000,000000

400000,000000

420000,000000

420000,000000

440000,000000

440000,000000

460000,000000

460000,000000 305000

0,00000

0

306000

0,00000

0

307000

0,00000

0

308000

0,00000

0

309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

Islas Canarias

!( Hydraena serricollis

10 0 105 km

!(

!(

!(

!(

420000,000000

420000,000000

430000,000000

430000,000000

440000,000000

440000,000000

450000,000000

450000,000000

460000,000000

460000,000000

470000,000000

470000,000000

480000,000000

480000,000000

490000,000000

490000,000000

500000,000000

500000,000000

307000

0,00000

0

307000

0,00000

0

308000

0,00000

0

308000

0,00000

0

309000

0,00000

0

309000

0,00000

0

310000

0,00000

0

310000

0,00000

0

311000

0,00000

0

311000

0,00000

0

312000

0,00000

0

312000

0,00000

0

Islas Canarias

!( Limnebius canariensis

10 0 105 km

!(

!(

!(

!(!(

!( !(

!(

!(

200000,000000

200000,000000

220000,000000

220000,000000

240000,000000

240000,000000

260000,000000

260000,000000

280000,000000

280000,000000

300000,000000

300000,000000

320000,000000

320000,000000

340000,000000

340000,000000

360000,000000

360000,000000

380000,000000

380000,000000

400000,000000

400000,000000

420000,000000

420000,000000

440000,000000

440000,000000

460000,000000

460000,000000 305000

0,00000

0

306000

0,00000

0

307000

0,00000

0

308000

0,00000

0

309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

Islas Canarias

!( Limnebius gracilipes

10 0 105 km

!(!(

!(

!(

!(!(

!(

!(!(

!( !(!(!(

!(!(!(

!(

!(!(!(

!(

!(!(

!(

!(

!(

!(

!(

!(

!(

!(!(

!(

!(

!(

!(

180000,000000

180000,000000

210000,000000

210000,000000

240000,000000

240000,000000

270000,000000

270000,000000

300000,000000

300000,000000

330000,000000

330000,000000

360000,000000

360000,000000

390000,000000

390000,000000

420000,000000

420000,000000

450000,000000

450000,000000

480000,000000

480000,000000

510000,000000

510000,000000

540000,000000

540000,000000

570000,000000

570000,000000

600000,000000

600000,000000

630000,000000

630000,000000 300000

0,00000

0

301000

0,00000

0

303000

0,00000

0

304000

0,00000

0

306000

0,00000

0

307000

0,00000

0

309000

0,00000

0

310000

0,00000

0

312000

0,00000

0

313000

0,00000

0

315000

0,00000

0

316000

0,00000

0

318000

0,00000

0

319000

0,00000

0

321000

0,00000

0

322000

0,00000

0

324000

0,00000

0

325000

0,00000

0

327000

0,00000

0

328000

0,00000

0

330000

0,00000

0

331000

0,00000

0

Islas Canarias

!( Ochthebius rugulosus

30 0 3015 km

!(

!(

200000,000000

200000,000000

210000,000000

210000,000000

220000,000000

220000,000000

230000,000000

230000,000000

240000,000000

240000,000000

250000,000000

250000,000000

260000,000000

260000,000000

270000,000000

270000,000000

315000

0,00000

0

315000

0,00000

0

316000

0,00000

0

316000

0,00000

0

317000

0,00000

0

317000

0,00000

0

318000

0,00000

0

318000

0,00000

0

319000

0,00000

0

319000

0,00000

0

Islas Canarias

!( Ochthebius heeri

4 0 42 km

!(

!(

!(

540000,000000

540000,000000

560000,000000

560000,000000

580000,000000

580000,000000

600000,000000

600000,000000

620000,000000

620000,000000

640000,000000

640000,000000

660000,000000

660000,000000

680000,000000

680000,000000

700000,000000

700000,000000

720000,000000

720000,000000

740000,000000

740000,000000 3100

000,00

0000311000

0,00000

0 312000

0,00000

0313000

0,00000

0

314000

0,00000

0315000

0,00000

0

316000

0,00000

0317000

0,00000

0 318000

0,00000

0

319000

0,00000

0 320000

0,00000

0

321000

0,00000

0 322000

0,00000

0323000

0,00000

0

324000

0,00000

0

Islas Canarias

!( Ochthebius bifoveolatus

10 0 105 km

!(

!(!(

!(

!(

!(

200000,000000

200000,000000

230000,000000

230000,000000

260000,000000

260000,000000

290000,000000

290000,000000

320000,000000

320000,000000

350000,000000

350000,000000

380000,000000

380000,000000

410000,000000

410000,000000

440000,000000

440000,000000

470000,000000

470000,000000

500000,000000

500000,000000

530000,000000

530000,000000

560000,000000

560000,000000

590000,000000

590000,000000

620000,000000

620000,000000

303000

0,00000

0

303000

0,00000

0

306000

0,00000

0

306000

0,00000

0

309000

0,00000

0

309000

0,00000

0

312000

0,00000

0

312000

0,00000

0

315000

0,00000

0

315000

0,00000

0

318000

0,00000

0

318000

0,00000

0

321000

0,00000

0

321000

0,00000

0

324000

0,00000

0

324000

0,00000

0

327000

0,00000

0

327000

0,00000

0

330000

0,00000

0

330000

0,00000

0

Islas Canarias

!( Ochthebius lapidicola

30 0 3015 km

!(!(!(

!(

!(

!(!(

!(!(

!(

!(

!(

!(

!(

!(

!(!(!(

!(!(

200000,000000

200000,000000

230000,000000

230000,000000

260000,000000

260000,000000

290000,000000

290000,000000

320000,000000

320000,000000

350000,000000

350000,000000

380000,000000

380000,000000

410000,000000

410000,000000

440000,000000

440000,000000

470000,000000

470000,000000

500000,000000

500000,000000

530000,000000

530000,000000

560000,000000

560000,000000

590000,000000

590000,000000

620000,000000

620000,00000030200

00,0000

00

303000

0,00000

0 305000

0,00000

0

306000

0,00000

0 308000

0,00000

0

309000

0,00000

0 311000

0,00000

0

312000

0,00000

0 314000

0,00000

0

315000

0,00000

0 317000

0,00000

0

318000

0,00000

0 320000

0,00000

0

321000

0,00000

0 323000

0,00000

0

324000

0,00000

0 326000

0,00000

0

327000

0,00000

0 329000

0,00000

0

330000

0,00000

0

Islas Canarias

!( Ochthebius quadrifoveolatus

20 0 2010 km

!(

600000,000000

600000,000000

610000,000000

610000,000000

620000,000000

620000,000000

630000,000000

630000,000000

640000,000000

640000,000000

650000,000000

650000,000000

660000,000000

660000,000000

670000,000000

670000,000000

680000,000000

680000,000000

690000,000000

690000,000000

700000,000000

700000,000000

319000

0,00000

0

319000

0,00000

0

320000

0,00000

0

320000

0,00000

0

321000

0,00000

0

321000

0,00000

0

322000

0,00000

0

322000

0,00000

0

323000

0,00000

0

323000

0,00000

0

324000

0,00000

0

324000

0,00000

0

325000

0,00000

0

325000

0,00000

0326

0000,00

0000

Islas Canarias

!( Ochthebius salinator

5 0 52,5 km

!(

!(

!(

!(

540000,000000

540000,000000

550000,000000

550000,000000

560000,000000

560000,000000

570000,000000

570000,000000

580000,000000

580000,000000

590000,000000

590000,000000

600000,000000

600000,000000

610000,000000

610000,000000

620000,000000

620000,000000

630000,000000

630000,000000

640000,000000

640000,000000

650000,000000

650000,000000

660000,000000

660000,000000

670000,000000

670000,000000

680000,000000

680000,000000

690000,000000

690000,000000

310000

0,00000

0

310000

0,00000

0311000

0,00000

0

311000

0,00000

0312000

0,00000

0

312000

0,00000

0313000

0,00000

0

313000

0,00000

0314000

0,00000

0

314000

0,00000

0315000

0,00000

0

315000

0,00000

0316000

0,00000

0

316000

0,00000

0317000

0,00000

0

317000

0,00000

0318000

0,00000

0

318000

0,00000

0319000

0,00000

0

319000

0,00000

0320000

0,00000

0

320000

0,00000

0

Islas Canarias

!( Ochthebius subpictus subpictus

8 0 84 km

!(

530000,000000

530000,000000

540000,000000

540000,000000

550000,000000

550000,000000

560000,000000

560000,000000

570000,000000

570000,000000

580000,000000

580000,000000

590000,000000

590000,000000

600000,000000

600000,000000

610000,000000

610000,000000

620000,000000

620000,000000

630000,000000

630000,000000

640000,000000

640000,000000

650000,000000

650000,000000

660000,000000

660000,000000

670000,000000

670000,000000

680000,000000

680000,000000

309000

0,00000

0

309000

0,00000

0310000

0,00000

0

310000

0,00000

0311000

0,00000

0

311000

0,00000

0312000

0,00000

0

312000

0,00000

0313000

0,00000

0

313000

0,00000

0314000

0,00000

0

314000

0,00000

0315000

0,00000

0

315000

0,00000

0316000

0,00000

0

316000

0,00000

0317000

0,00000

0

317000

0,00000

0318000

0,00000

0

318000

0,00000

0319000

0,00000

0

319000

0,00000

0320000

0,00000

0

320000

0,00000

0

Islas Canarias

!( Ochthebius tivelunus

8 0 84 km

!(!(

!(!(!(!(

!(

!(

!( !(

!(

!(

!(!(

!(!(!( !(

!(

!(

!(!(

!( !(!( !(

190000,000000

190000,000000

210000,000000

210000,000000

230000,000000

230000,000000

250000,000000

250000,000000

270000,000000

270000,000000

290000,000000

290000,000000

310000,000000

310000,000000

330000,000000

330000,000000

350000,000000

350000,000000

370000,000000

370000,000000

390000,000000

390000,000000

410000,000000

410000,000000

430000,000000

430000,000000

450000,000000

450000,000000

470000,000000

470000,000000

490000,000000

490000,000000

510000,000000

510000,000000

530000,000000

530000,000000 304000

0,00000

0

305000

0,00000

0

306000

0,00000

0

307000

0,00000

0

308000

0,00000

0

309000

0,00000

0

310000

0,00000

0

311000

0,00000

0

312000

0,00000

0

313000

0,00000

0

314000

0,00000

0

315000

0,00000

0

316000

0,00000

0

317000

0,00000

0

318000

0,00000

0

319000

0,00000

0

320000

0,00000

0

321000

0,00000

0

322000

0,00000

0

323000

0,00000

0

324000

0,00000

0

325000

0,00000

0

326000

0,00000

0

327000

0,00000

0

328000

0,00000

0

Islas Canarias

!( Dryops gracilis

50 0 5025 km

Resumen

365

LOS COLEPTEROS ACUTICOS DE LAS ISLAS CANARIAS

Resumen

Este estudio se ha realizado en las siete islas del archipilago canario. En esta zona la disponibilidad de agua es

escasa, sin embargo algunos de los ecosistemas acuticos presentes son nicos en un contexto europeo. A pesar de su

enorme inters de conservacin, se ha observado que se encuentran en regresin en las ltimas dcadas, debido

principalmente a la contaminacin y canalizacin.

En las islas Canarias estn catalogadas 53 especies de colepteros acuticos, de las cuales 14 especies y una

subespecie son endemismos canarios.

En las islas Canarias, desde el siglo XIX, se ha observado una marcada ralentizacin en el incremento de

especies registradas, lo que puede ser interpretado como que actualmente existe una aproximacin importante hacia el

nmero mximo de especies que pueden albergar. Por lo tanto, este estudio se ha centrado en complementar la

distribucin inter e intrainsular de las especies, ampliando la distribucin interinsular de 10 de ellas.

Se ha evaluado la vulnerabilidad de las especies de colepteros acuticos de las islas Canarias a tres escalas:

regional, nacional e internacional. Las especies con mayor vulnerabilidad a escala regional son Limnebius similis, L.

punctatus, L. canariensis, Hydroporus compunctus, H. pilosus y Agabus conspersus y a escala nacional e

internacional son Limnebius similis, L. punctatus, L. canariensis, Hydroporus compunctus e H. pilosus

Adems se identifican los enclaves acuticos con mayor biodiversidad de las islas Canarias y por lo tanto

aquellos con mayor inters de conservacin. Para ello se ha utilizado como herramienta a los colepteros acuticos y

se ha aplicado un algoritmo iterativo de complementariedad.

La actual Red de Espacios Naturales Protegidos en complementacin con la Red Natura 2000, resulta

suficiente y adecuada para la conservacin de los ecosistemas acuticos de las islas Canarias, ya que la gran mayora

de los principales ecosistemas acuticos de las 10 cuadrculas seleccionadas como reas prioritarias de conservacin

quedan incluidos, al menos en parte, dentro de algn tipo de espacio protegido.

THE WATER BEETLES OF THE CANARY ISLANDS

Abstract

This study was carried out in the Canary Islands. In this area, the water availability is limited, however some of these

aquatic ecosystems are unique in a European context. In spite of their huge convervational interest, in recent decades

it has been observed that they are in declive, main due to contamination and channeling.

A total of 53 species of aquatic Coleoptera from the Canary Islands were recorded, 14 species and one

subspecie are Canarian endemics.

Biodiversity of water beetles in the Canary Islands is known but not its distribution in the different islands

and within each island. This study helps to improve the knowledge to the distribution of species of aquatic

Coleoptera, extending the insular distribution of 10 species.

The water beetles species from the Canary Islands were ranked according to their conservation priority or

their vulnerability to extinction at three scales: local, national and international. Limnebius similis, L. punctatus, L.

canariensis, Hydroporus compunctus, H. pilosus and Agabus conspersus have the highest degree of vulnerability al

local level, and Limnebius similis, L. punctatus, L. canariensis, Hydroporus compunctus and H. pilosus at national

and international levels.

The aquatic ecosystems with the highest biodiversity, and therefore the highest conservational interest, in

the Canary Islands are identified. For that purpose, the water beetles were used as indicators in an iterative algorithm

of complementarity.

The current network of Protected Natural Areas along with the net Natura 2000 are sufficient for the conservation of aquatic ecosystems of the Canary Islands, as they are included the ten 10x10 km squares containing

the aquatic ecosystems with the highest biodiversity, at least in part.

Documents

TESIS JOSE GUTIERREZ PARTE 2.pdf